The NHBS Guide to UK Caterpillar Identification

Caterpillars are part of the life cycle of moths or butterflies which is known as complete metamorphosis. This life cycle includes four stages: egg, caterpillar (also known as the larval or feeding stage), pupa (the transition stage) and adult (the reproductive phase). With over 2,600 species of moth and 60 species of butterfly in the British Isles, there are a large variety of caterpillars present in our countryside.

There are several stages of caterpillar growth called instars, during which the caterpillar sheds its skin as it grows. Colouration, size and patternation can vary between these instars. Additionally, species can have different variations of caterpillars, including different colour forms. Several species are listed below, grouped by key characteristics such as colour, patternation and features.

Hairy caterpillars

There are many ecological functions of hair-like structures on caterpillars including defence and camouflage. These hairs, called setea, can be almost invisible to the naked eye, while others make them easier to see. Two types of caterpillar hair can cause harm to humans and pets: urticating, which are itchy, non-venomous hairs that can irritate the skin, and stinging hairs, which are hollow spines that have poison-secreting cells that can cause a range of health issues if they enter the skin.  

 

A black, yellow and red spotted caterpillar with several tufts of orange and brown hair
Image by Kjetil Fjelleim via Flickr

Knot Grass moth (Acronicta rumicis): Colour can vary between light gingery brown to near black, with patches of rusty brown hair and a broken line of white dorsal patches. They also have a wavy white line on their sides, broken with bright orange/red spots. They grow up to 40mm in length. Can be confused with the caterpillars of Brown-tail and Yellow-tail moths. Foodplants include Knot Grass as well as Broad-leaved Dock, plantains, Bramble, Hawthorn, Common Sorrel, heather, and Purple Loosestrife.

 

A fluffy black caterpillar with orange stripes
Image by Gail Hampshire via Flickr

Fox Moth (Macrothylacia rubi): Very hairy, up to 70mm long, dark brown with an orangey stripe down the length of its body. Caterpillars in earlier stages of development may have distinctive orange or yellow bands. Commonly feeds on heathers, Bilberry, Creeping Willow, Bramble, Meadowsweet and Salad Burnet.

 

a caterpillar on a leaf. It is very fluffy with long black hairs on top and shorter orange hairs on its underside
Image by Dean Morley via Flickr

Garden Tiger moth (Arctia caja): Also known as the woolly bear caterpillar due its very long hairs. Grows up to 55mm long and has a dark red dorsal area with white tipped hairs,an orangey red underside, and small white markings along its sides. Feeds on a variety of herbacious and garden plants including Common Nettle, Broad-leaved Dock, burdocks and Hound’s-tongue. 

 

A dark brown caterpillar with long, spiking hairs and white and orange parallel patches running along its back
Image by Chris Cooper via Flickr

Brown-tail moth (Euproctis chrysorrhoea): Can measure up to 30mm long, black with white markings down its sides and two distinctive orangey red ‘warts’ on its back near its tail. Be aware that its hairs are toxic to humans. Feeds on plants in the Rosaceae family including Hawthorn, Blackthorn, Plum, Cherry, Rose and Bramble.

 

A large grey caterpillar covered in long white wispy hairs
Image by Janet Graham via Flickr

Miller moth (Acronicta leporina): Up to 35mm long with very long white or yellow hairs that swirl to one side. The body is often a pale green to brown depending on the development stage but this can be hard to see under the hairs. Usually found on birch or Alder trees. 

 

A lime green moth with tufted white bristly hair
Image by gailhampshire via Flickr

Pale Tussock moth (Calliteara pudibunda): Greenish yellow hairs with a black body showing through in bands between tufts. The hairs can vary in colour and can be white, brown or pink. They also have a tail tuft that varies in colour but is usually brown, pink or red. This can be absent in some individuals. The four, tussocky tufts on their dorsal are frequently white, brown or yellow. Feeds on a variety of broadleaved trees and shrubs including Hawthorn, Blackthorn, Crab Apple, oaks, birches and Hazel.

 

A yellow fluffy moth with white spots down the back and orange tufts
Image by Jon Brinn via Flickr

Sycamore moth (Acronicta aceris): Up to 40mm long with thick hair that is either yellow, brown or orange . They have bold white spots down their back, outlined in black, as well as tufts of dark orange or bright red hair on their back. Foodplants are most commonly Sycamore, Field Maple and Horse-chestnut.

 

A fluffy black caterpillar with a single brown line down the back
Image by Odd Wellies via Flickr

White Ermine moth (Spilosoma lubricipeda): Approximately 40mm long with a red, orange or pale dorsal line. Caterpillars at later development stages are covered in spines that can be reddish brown, dark brown or even black.

 

Brown caterpillars
A black moth with a purplish tint and a large head. it is hairless
Image by Aah-Yeah via Flickr

Elephant Hawk-moth (Deilephila elpenor): Thick bodies that grow up to 8cm in length, usually dark brown but bright green forms also occur. The name derives from their smaller, trunk-like head that extends from its more bulbous neck. They feature a spiked tail and four eyespots, although the second pair can be less visible on darker individuals. Most frequently found on Rosebay Willowherb, Great Willowherb, other willowherbs and bedstraws. 

 

A striped brown caterpillar
Image by David Short via Flickr

Square-spot Rustic moth (Xestia xanthographa): Greenish ochre in colour, with pale lines on its back and edged with dark, long, slanted markings on its sides in a row. Mainly feeds on grasses, plantains and Cleavers.

 

A pale brown caterpillar with darker brown stripes and dark brown spots
Image by Bjorn S via Flickr

Large Yellow Underwing moth (Noctua pronuba): Grows to a length of 45–50mm. Its body can be various shades of brown and green, with three lines down its back and dark patches on the inner side of the outer two lines – similar to the Square-spot Rustic. They also have darker sides with a lighter stripe above the legs. Feeds on a wide range of herbaceous plants and grasses including docks, brassicas, marigolds and Foxglove.

 

A hairless brown moth with speckles of cream and pink, and a cream stripe down its back
Image by Janet Graham via Flickr

Dot Moth (Melanchra persicariae): These caterpillars can reach up to 45mm in length and can be different shades of brown and green. They have three pale, distinctive lines on the dark prothoracic plate behind their head, as well as dark and light chevrons along a pale dorsal line down their backs. Feeds on a wide range of herbaceous and woody plants including Common Nettle, White Clover, Ivy, Hazel, Elder and willows.

Many of these caterpillars can also have a green form.   

Black and yellow/orange patterned caterpillars
Two black and lime green spotted caterpillars, with light coloured sparse hairs over the body
Image by Katja Schulz via Flickr

Large White butterfly (Pieris brassicae): Pale green-yellow in colour with black spots along its body. Visibly hairy. Also known as a Cabbage White due to its preference for cabbages as a food plant. 

 

A black moth with yellow and orange stripes with fine, long white hairs
Image by Dean Morley via Flickr

Buff-tip moth (Phalera bucephala): Distinctive caterpillar with a trellised black and yellow patterning and covering of pale hairs. The face is black and has an inverted yellow V. When fully grown this caterpillar measures up to 75mm in length. Most frequently found on sallows, birches, oaks and Hazel. 

 

A yellow-green caterpillar with black spots and stripes and short white hairs
Image by Peter O’Connor via Flickr

Six-spot Burnet moth (Zygaena filipendulae): Caterpillars feature a series of yellow and black dots on a green or greenish-yellow body. Feeds on Common Bird’s-foot-trefoil or occasionally Greater Bird’s-foot-trefoil. 

 

Dark brown/black caterpillars on a leaf. They have two cream stripes running the length of their back and have multiple protruding hairs
Image by Gilles San Martin via Flickr

Small Tortoiseshell butterfly (Aglais urticae): Caterpillars are initially black but show increasing variation in colour, with many developing pale yellow lines down their back and sides (some, however, may remain pure black). They have small clusters of short yellow spines and are fully grown at 30mm. Usually found on Common Nettle leaves.

 

A pale green caterpillar is on a plant. It has yellow and black spots in a repeating pattern
Image by Amanda Slater via Flickr

Mullein moth (Curcullia verbasci): One of the most striking and distinctive caterpillars to be found in Britain, they have a mixture of repeating black and yellow markings on a pale bluish-grey body. When fully grown they measure almost 50mm in length. Foodplants include mulleins, Common Figwort, Water Figwort and buddleias.

 

A green caterpillar with a black head. it has brown and green stripes down the length of its body and black spots
Image by hedera.baltica via Flickr

Box Tree moth (Cydalima perspectalis): Box Tree moths were introduced accidentally from south-east Asia and are a pest of Box trees. Caterpillars have green and black stripes running the length of the body, and the head is shiny black. Each of the body segments has white hairs and eyelike markings. 

 

 Black and spiky caterpillars
A black, spiked caterpillar with small white spots
Image by Bernard Dupont via Flickr

Peacock butterfly (Aglais io): Unlike the brightly coloured adult Peacock butterfly, the Peacock caterpillar has a velvety black body with small white spots and short spines on each segment. Most commonly feeds on Common Nettle and Hops.

 

A caterpillar with spikes and small white hairs. It has a black upper and orange lower with cream stripes
Image by Dean Morley via Flickr

Painted Lady butterfly (Vanessa cardui): Often found on thistles, Painted Lady caterpillars live for 5-10 days in a loosely woven silk nest inside which they feed continuously. They have dark bodies with pale narrow yellow-cream stripes. Particularly on younger larvae, spines can be alternating light and dark.

 

A group of black spiky caterpillars spinning a web on a green stalk
Image by Gilles San Martin via Flickr

Marsh Fritillary butterfly (Euphydryas aurinia): Caterpillars are black and hairy and initially live in groups on a larval web which is woven on the bottom-most leaves of Devil’s Bit Scabious plants. Prior to pupation, at the end of April, caterpillars will finally disperse to live independently. 

 

A brown caterpillar with white speckles and rings of black spikes
Image by Tim Worfolk via Flickr

Red Admiral butterfly (Vanessa atalanta): Caterpillars are black and spiny with a yellow stripe down each side and fine hairs along the body. They can be tricky to spot as they use silk to bind nettle leaves together to make a protective tent inside which they feed. 

 

Green caterpillars
A large green caterpillar on a tree trunk. it has small yellow spots and stripes on its back and two horns at one end
Image by Odd Wellies via Flickr

Lime Hawk-moth (Mimas tiliae): Caterpillars are distinctive having a large green body with pale yellow streaks on each segment and a bluish ‘horn’ at the tail end. Turns purple a short time before pupation. Foodplants include Limes, elms, Downy Birch, Silver Birch and Elder. 

 

A chunky green caterpillar. it has pink stripes on its legs and small green spikes on its body
Image by Patrick Clement via Flickr

Poplar Hawk-moth (Laothoe populi): A thick and chunky, bright green caterpillar with faint yellow lines running diagonally along the body. The tail end has a yellow ‘horn’ and some individuals have small, dark spots. Food plants include poplars, sallows and willows.

 

A chunky light green moth, it has a brown and yellow horn at one end and diagonal stripes along its back
Image by Jo Garbutt via Flickr

Privet Hawk-moth (Sphinx ligustri): Bright, lime-green caterpillar with white and purple stripes and a pale yellow spot on each segment. The tail end has a black curved hook. Usually found on Wild and Garden Privets, Ash, Lilac and Guelder-rose.

 

A round lime green caterpillar is walking along a plant stem. It is covered in very small white dots and subtle white stripes
Image by Julian Smith via Flickr

Eyed Hawk-moth (Smerinthus ocellata): Closely resembles the Poplar Hawk-moth caterpillar in that it is bright green with diagonal yellow lines. When mature it can be distinguished by its bluish tail horn. Foodplants include Apple, willows and sallows. 

 

a long, thin caterpillar with light green stripes running the length of its body
Image by Dean Morley via Flickr

Speckled Wood butterfly (Pararge aegeria): Bright green with faint dark green and yellow stripes running longitudinally along the length of the body. Feeds on False Brome, Cock’s-foot, Yorkshire-fog and Common Couch.

 

A green moth with a brown stripe down its back. It has a black hook at the end of its body and yellow patches along the side of its body
Image by Aah-Yeah via Flickr

Pine Hawk-moth (Sphinx pinastri): Dark green caterpillar with a brown stripe along the centre of its back and cream dashes that run either side of this. It has a brown head and a black tail horn. Feeds mainly on Scots Pine.

 

A lime green caterpillar with light black speckling and a yellow line running the length of the flank
Image by Ben Sale via Flickr

Bright-line Brown-eye moth (Lacanobia oleracea): Green caterpillar with a bright yellow line along its sides and tiny black spots. Found on a variety of herbacious and woody plants such as Common Nettle, Fat-hen, willowherbs, Hazel and Hop. Sometimes a pest of cultivated Tomatoes.

 

A slender green moth with rings of white spots. It has two white stripes running along its back leading toward a black hook at the base of the body
Image by Lies Van Rompaey via Flickr

Hummingbird Hawk-moth (Macroglossum stellatarum): Caterpillars are mainly green and have a thick, cream-yellow stripe running along the sides with a white line above. The tail horn is black with a yellow tip when mature. Feeds on Lady’s Bedstraw, Hedge Bedstraw and Wild Madder.

 

A green caterpillar with sparse dark hairs and two white stripes running the length of the back
Image by Mick Talbot via Flickr

Straw Dot moth (Rivula sericealis): Green caterpillar with two cream stripes running along the back creating a repeating hourglass pattern between them. Covered in long fine hairs. Not often seen, the caterpillars feed on a variety of grass species.

 

A plump green caterpillar with sparse hairs
Image by Artur Rydzewski via Flickr

Silver Y moth (Autographa gamma): Relatively easy to identify as it has only two sets of prolegs (small fleshy stubs beneath the body) and a rear clasper which means it walks with an arched body. It has a green body with a series of white wavy lines which may be broken by pale circles in later instars. Feeds on a range of low-lying herbacious plants including bedstraws, clovers, Common Nettle, Garden Pea and Cabbage. 

 

A plump green caterpillar with black speckles on its underside
Image by Harald Supfle.

Kentish Glory moth (Endromis versicolora): Large green caterpillar with diagonal pale stripes on each segment. Usually found on Silver Birch and less often on Downy Birch and Alder.

 

A chunky green caterpillar with orange spots. the spots are covered in spikes and small fine hairs
Image by Odd Wellies via Flickr

Emperor Moth (Saturnia pavonia): Green with black hoops containing yellow wartlike spots. Common in scrubby places whether they often feed on heathers, Meadowsweet, Bramble, Hawthorn and Blackthorn, amongst others. 

 

A vibrant green caterpillar with sparse white hairs
Image by gailhampshire via Flickr

Angle Shades moth (Phlogophora meticulosa): Usually green but can be mixed with shades of brown and/or yellow. A fine pale line runs down the back and a pale band runs down the sides of the body. Foodplants include a range of herbaceous and woody plants such as Common Nettle, Hop, Red Valerian, Bramble and Broad-leaved Dock. 

 

 Others
A plump palegreen caterpillar with black bands across its back and orange spots
Image by Frank Vassen via Flickr

Swallowtail butterfly (Papilio machaon): Striking bright green caterpillar with black bands and orange spots. British Swallowtail caterpillars feed solely on Milk-parsley. 

 

A black and orange striped moth on ragwort flowers
Image by Smudge 9000 via Flickr

Cinnabar moth (Tyria jacobaeae): Easy to identify having bold gold and black stripes. Most commonly feeds on the leaves and flowers of Common Ragwort where they can be found in their hundreds.

 

A brown caterpillar with thick spikes
Image by Darius Bauzys via Flickr

High Brown Fritillary butterfly (Argynnis adippe): Black caterpillar with a checkered pale pattern and yellow/buff spines. Covered in fine black bristles. Feeds on Common Dog-violet and Hairy Violet. 

 

A yellow caterpillar covered with black spots and patches. it has orange stripes running down the side of its body
Image by Conall via Flickr

Magpie moth (Abraxas grossulariata): Distinctive caterpillar with a creamy-white body, rows of black and white spots and an orange stripe that runs along the length of the body on the lower sides. Feeds on a range of deciduous trees such as Blackthorn, Hawthorn and Hazel as well as currant and gooseberry bushes. 

 

A short, rounded caterpillar (shaped like a woodlouse) with fine pink hairs. It is pink in colour with a green stripe running on either side of its body
Image by Gilles San Martin via Flickr

Small Copper butterfly (Lycaena phlaeas): Slug-shaped caterpillar covered in tiny white hairs. Exists in two forms: a purely green form and a green and pink striped form. Main foodplants are Common Sorrel and Sheep’s Sorrel. 

 

A black caterpillar with orange spots and stripes. the spots have spikes growing out of them that are also orange in colour
Image by Gilles San Martin via Flickr

Comma butterfly (Polygonia c-album): Mainly coloured brown and black with a large white mark towards the rear end of its back. Preferred foodplant is Common Nettle. 

 

A black caterpillar with an orange stripe running centrally along the back. On its sides are white patches with long white hairs growing out of them, the rest of the body has long black hairs
Image by gailhampshire via Flickr

Yellow-tail moth (Euproctis similis): Black caterpillar with a small hump behind its head. Two red/orange lines run along the back with a row of white markings wither side of them. They are covered in long black hairs and shorter white ones. Feeds on a wide selection of broadleaf trees and shrubs including Hawthorn, Blackthorn, oaks, roses, Hazel and willows.

 

A group of caterpillars. They are orange and blue striped with orange hairs protruding from the sides of its body
Image by gailhampshire via Flickr

Lackey moth (Malacosoma neustria): Large orange, blue and white striped caterpillars that are covered with fine orange hairs. Often feed in large groups on broadleaved trees and shrubs including Blackthorn, Hawthorn, cherries, Plum and Apple.

 

Recommended Reading:  
Field Guide to the Caterpillars of Great Britain and Ireland

This Bloomsbury Wildlife Guide allows identification of the common moth and butterfly caterpillars of the British Isles.

 

 

A Guide to the Caterpillars of the Butterflies of Britain and Ireland

A second edition fold-out guide to 57 of Britain’s butterfly species.

Equipment in Focus: Royal Entomological Society Bug Hunting Kits

Shows the bug kit- containing a net, ID guide, pooter and collecting pots

The Royal Entomological Society (RES) is an organisation dedicated to advancing the field of insect science. Through encouraging open communication, research and publication, the RES hopes to enrich the world with entomology 

Developed in collaboration with the RES, the Royal Entomological Society Bug Hunting Kits provide naturalist users with the tools to safely capture, observe and identify British insects. Kitted with sweep nets, collecting pots and a pooter to capture your insects, you will also be provided with a hand lens, ID guide and optional forceps for identification of species you find. 

Suited for aspiring entomologists, The Royal Entomological Society Educational Bug Hunting Kit includes a copy of A Naturalists Guide to The Insects of Britain and Northern Europe. The Royal Entomological Society Advanced Bug Hunting Kit provides a technical alternative for more experienced naturalists, with additional pointed forceps and the Collins Complete Guide to British Insects. Here we take a closer look at what’s included in these exclusive kits.  


A hand holding a net, sweeping in a bush of nettles.

As seen on Countryfile, the Standard Sweep Net provides users with a simple, lightweight (280g) net for catching invertebrates. The short, 15cm handle has a foam grip for improved control and a lightweight aluminium frame. The net itself is made in the UK and features a soft calico bag attached via Velcro to the frame, making it easy to remove for washing.

 

A hand holding a pooter- a plastic chamber with two long plastic tubes used to entrap invertebrates

At the core of this kit is the NHBS Insect Pooter. Expertly designed and manufactured at our facilities in Devon, this piece of kit can safely capture a wide range of invertebrates. Affordable and simple to use, this item allows the user to observe specimens in a see-through chamber. The chamber is topped with a 2.5× magnification lens for easy viewing and identification. The pooters components can be removed and cleaned for sanitation between sampling.  

 

An alder fly in a collecting pot on a page of an identification guide showing species of flying insects

Each kit comes with five 60ml Collecting Pots for specimen handling and collection. The collecting pots have secure screw-on lids, made with see-through polypropylene for easy, clear viewing.  

 

A hand holding a magnifying hand lens over a ladybird on a leaf.

The handy Double Loupe Hand Lens provided with this kit is only 30mm in diameter, comprising two silicate glass lenses, 5× and 10× magnification. The lenses of this sturdy pocket magnifier fold into a protective casing, keeping them clear from scratches between use. Lightweight and compact, this hand lens is highly portable and is ideal for people of all ages. 

 

a pair of metal forceps with a beetle on a muddy tree stump

Made from a non-magnetic stainless-steel alloy, the Super Fine Pointed Forceps are manufactured with fine points for precision use. Included with the Advanced Bug Hunting Kit, these precise forceps are not serrated to minimise damage to delicate specimens, and at 11cm are a handy size for transportation and use in-field.  

 

Front cover of the Collins Complete Guide to British Insects

Provided with the Advanced Bug Hunting Kit, the Collins Complete Guide to British Insects is a photographic field guide to common and unusual insect species across Britain. This extensive work covers over 1,500 species, providing descriptions and detailing where, and when, to observe them. With detailed photographs for each species, differences between similar organisms are highlighted to aid identification. This book covers a range of insects, from bugs and bees to moths and mayflies.  

 

Front cover of a Naturalists guide to the insects of britain and northern europe

A Naturalists Guide to the Insects of Britain and Northern Europe is provided with the Educational Bug Hunting Kit. This easy-to-use ID guide is ideal for nonspecialist naturalists, with high quality photos of over 280 insect species. A description of appearance, associated habitats, habits and conservation status are outlined for each species. The guide also includes life cycles and describes the conservation of the group.     

 


The mission of The Royal Entomological Society is to enrich the world with insect science- doing this through events, books and supporting young people in gaining skills in entomology.

The Royal Entomological Society receives 10% from the sale of this kit to support their cause.

Spring Exploring: Equipment for Wildlife Watching

As spring emerges, naturalists across the UK are dusting off their kit to begin exploring the great outdoors. From bird watching to bug hunting, we have equipment to help you explore. Below, we have compiled a list of must-have equipment for wildlife watching this spring. 


Viking Cygnus Monocular 

Young boy looking through a monocular in the woods.
The Viking Cygnus Monocular in-field.

Excellent optics combined with a grippy, rubberised armour make this handy monocular the ideal companion this spring. At only 287g, The Viking Cygnus Monocular is a lightweight, showerproof monocular with a small form factor, designed for easy handling for any hand size. Created for quick access and target acquisition, this monocular features a smooth action barrel adjuster for precise focus adjustment and a larger objective lens for high colour imagery.  

For a smaller, even more lightweight alternative weighing only 140g, the 8×25 MK2 magnification monocular is ideal for quick and easy use, where a smaller objective diameter lens is counteracted by its handy size. 

Opticron Explorer Compact Binoculars 

Black binoculars.

The Explorer Compact Binoculars by Opticron would make an excellent addition to any naturalists kit this spring. A fully armoured, roof prism body provides comfort and extra grip for comfortable carrying infield. With a weatherproof, fold-down design, these are ideal for transport, and can be stored easily due to their size and weight (195g). The ribbed focus wheel and twist-type eye cups ensure a good field of view with easy focusing, and the use of multicoloured lens and high reflection coated prisms provide bright, crisp images. Available in 8 x 21 and 10 x 21.  

Nikon Sportstar EX DCF Compact Binoculars 

Black binoculars

Available in 8×25 and 10×25, these high-quality, pocket-sized binoculars are waterproof and fog free. Turn and slide rubber eye cups allow for easy positioning, and multilayer coated lenses deliver a high optical performance with great clarity and well-balanced colour. The field of view is ideal for observing large landscapes, and partnered with a good close focusing distance, these compact binoculars also work great with insects. Weighing only 300g, these ultra-lightweight binoculars are ideal for travelling or working infield.  

Crushable Pocket Butterfly Net

A hand holding a butterfly net

Designed with a spring steel frame, this Crushable Pocket Butterfly net can twist for an easy collapse and can be folded down to pocket-size. This foldable, yet robust design allows for easy transportation and storage whilst in-field. The net is supplied with a short, brass handle but can also be used with telescopic and push-fit net handles if you wish to extend its reach.  

Walkstool Basic 

A black and grey folding stool.

Designed and manufactured in Sweden, the Walkstool Basic is a simple, 3-point stool made for outdoorsmen of any kind. Suitable for home use and in-field work, this compact resting stool weighs only 725g, making it ideal for packing and transporting. Available in 24”, the Walkstool Basic is designed with comfort and sturdy support in mind. This highly portable stool has plastic foot ends and telescopic, extendable legs to account for uneven terrain. The sturdy aluminium frame and durable polyester seat make this stool a worthy addition to any explorers kit this spring.  

Pocket Guide to the Bumblebees of Great Britain and Ireland 

A hand holding a small book with a bumblebee on the front.

This handy, pocket-sized guide to the naturally occurring bumblebees of Great Britain and Ireland is a richly illustrated work accessible to beginners and more experienced naturalists alike. Each species has a dedicated double page spread, detailing its characteristics, habitat, distribution and sex differentiation, among others. This portable pocket guide provides an ‘at-a-glance’ guide to species. Ideal for exploring this spring, this handy book provides an informative peak into the world of bumblebee identification.  

Pocket Guide to the Butterflies of Great Britain and Ireland  

A hand holding a guide to butterflies with a background of grass

Another publication in the Bloomsbury Wildlife Guides collection, the Pocket Guide to the Butterflies of Great Britain and Ireland features over 600 detailed illustrations on each species and their life stages. Detailing species information, distribution and life history, this pocketbook provides an accessible, easytouse guide to butterflies in Britain.   

Field Studies Council Fold-out Guides  

Identification guide showing seaweed species

The Field Studies Council Fold-out Guides are ideal for days where full-size field guides are cumbersome. These handy species identification charts cover an eclectic range of themes, from mammal tracks and woodland plants to seashells and jellyfish. These weatherproof guides are a practical accompaniment to a spring stroll, find the full collection here 

No Mow May: A Celebration of Wildflower Power

This spring, traditional British lawns are out. Throughout the month of May, Plantlife urges us to let our gardens be wild with #NoMowMay. This exciting initiative encourages us to embrace a wild lawn this spring, providing plants, invertebrates and other wildlife the opportunity to make our gardens a home. No Mow May could transform your green spaces into a colourful kaleidoscope of flowers you never knew were there. From buttercups to bee orchids, here at NHBS we have had an astonishing array of wildflowers in previous years, and we are hoping that this year will be the same!

Knowing when, and how, to mow your lawn to encourage wildflower growth and minimise grass domination can be confusing, and there is no ‘one size fits all’ answer to supporting native wildlife. In anticipation of May, we outline the important things to consider when maintaining your lawn over the coming seasons.


Tightly manicured garden lawns are unable to host the diverse communities associated with a natural space. The artificially constructed environment, with uniform grass length and limited species, prevents our native wildflowers from blooming and our vital insects from settling. Lawn feeds and fertilisers often used to maintain our lawns can result in unnaturally high levels of soil fertility. Such levels can unintentionally diminish the diversity of flora within our gardens, since native wildflowers are adapted to low-nutrient conditions. Associated with higher carbon emissions, time consumption and overall cost, many are steering clear of a high maintenance lawn this spring. 

A spring-flowering lawn provides a whole host of benefits for the wildlife within our gardens. Opting for a wild, native lawn provides essential breeding habitats, food sources and physical protection for a number of species. These spaces give wildflowers a chance to bloom and set seed, benefitting both insects, and the predators who rely on them.  

 

A bee orchid in the centre, in front of a wild lawn
Our Bee Orchid (Ophrys apifera) from #NoMowMay 2022. Image by Oli Haines.

So, how and when should we mow?   

Less is more! Switching up your mowing routine, or refraining from a mow in some areas, is a great way to maximise diversity in your garden. After a short time, your outdoor spaces can flourish into a haven for wildlife. From voles to vetches, and even British reptiles, watch your garden transform from monoculture to a wild refuge.  

Varied grass length, wild edges, or longer patches of lawn are great for attracting local wildlife to your garden. You may find orchids, ox-eye daisy and knapweed in these longer areas, which also provide cover for small mammals that may be wandering through, and shorter areas can boost pollen availability from low-lying flowers, like buttercups and clover. Plantlife advocates for a varied mowing approach with longer patches throughout the garden, alongside shorter areas (aiming to mimic grazing pressures of different herbivorous species in the wild). For instance, you might decide to maintain shorter pathways and areas around patios, but allow other areas of your green spaces to grow freely.  

It is important to remove cuttings after lawn maintenance to prevent excess nitrogen in the soil, thus reducing nitrophilic plants (species with a preference for nitrate rich habitat, typically from fertilisers and the decomposition of organic material) in your garden. ‘Cut and rot’ management can be counterproductive when cultivating wildflowers, as low levels of soil nutrition are preferred by many and will harbour the most diversity. In fact, frequent fertilisation and additional nutrition can result in an overall decline of wildflowers, leading to a dominance of nitrophilic plant species.   

A garden during No Mow May with varied grass length, wildlife corridors and vegetable patches.
A garden with varied grass length during No Mow May. Image by Allan Harris via Flickr.

Knowing when, and how, to mow during the year is key to maximise flowering of wildflower species, while simultaneously preventing grass domination: to do this, it is generally recommended to mow three times a year; early spring, late summer and in autumn.  

A 3-inch, early spring mow is beneficial to kickstart the season, promoting early growth and blooming.  An early mow can also help to tackle nitrophiles, like nettles and cow parsley. This can help to prevent competition, allowing wildflowers to grow undisturbed. However, be wary of mowing too early, as this can prevent wildflower seeding and will impact your gardens growth next year.  

A summer mow in late July, or August, removes the previous growth, encouraging the bloom of wildflowers later in the season. As far as insects are concerned, the later the mow, the better. Insect species tend to hatch in the warmer parts of spring and summer, so a mow in late August will prevent harm to hatching individuals. 

Around late November, an autumn mow can help to promote reseeding and encourages germination in the following spring. Allow the wildflowers in your lawn to finish flowering and let them go to seed, a mow after this allows the seedheads to disperse seeds into your lawn. An autumn cut can also keep grass growth under control, further encouraging germination.  

There are also certain considerations to be wary of when forming wild areas in your garden. These habitats will attract a great number of species, who may make your lawn a home. Best practice involves leaving an area of your lawn untouched to house these species, but if you are looking to tidy up your garden after No Mow May, wildlife must be considered. Wildlife in our lawns can be harmed in the process of tidying up our outside spaces. It is recommended to disturb, or walk through patches to be maintained to shoo species from the area. On the first mow, start with a higher cut to give smaller animals a chance to escape. When mowing the lawn, start with garden paths and areas of high footfall, working toward the edges of the garden. This, again, provides wildlife with an escape route through the boundaries of your garden. If your garden has fences or hedgerows, a wildlife corridor along your borders is another way to support visiting animals. Untouched, or lightly managed, strips along these areas can provide a safe space for travel around the garden, providing cover and protection from predators.  

hedgehog looking out from a bush
Hedgehog by Kalle Gustafsson via Flickr.

How can we prepare for No Mow May?  

If you currently use fertilisers, lawn feed, moss killers or pesticides, abandoning the use of these additives in your garden will allow the soil to recover from these harmful chemicals. This can provide microscopic and invertebrate soil communities a chance to recover, improving the overall health of your soil.  

For some of us, early bloomers may already be present in our gardens. Cowslip, violets and primroses may be popping up on our lawns, showcasing the first few flowers of the season. You may consider allowing these to go undisturbed, giving them a head start for spring. Having said that, the best way to prepare for No Mow May is a 3-inch April cut to encourage a strong period of spring growth.  

Whether or not you decide to mow the lawn this spring, consider leaving an area of your garden wild. Whether this be a natural lawn or rough borders, we hope you feel inspired to take part in this year’s #NoMowMay! 

 

The NHBS Guide to UK Snail Identification

Snails are a common feature in our gardens and parks. You may have particularly noticed them if you have a vegetable or plant patch, as they feed on the leaves, flowers and fruits of many of our food plants. There are over 40,000 species of land snail, although only around 120 occur in Britain.  

There are several useful features for identifying the correct species. The overall shape, in terms of the ratio of height to breadth, is important, as species can vary between a wide, round, flattened shape to tall and thin. The shape, colouration and thickness of the mouth of the shell can often be used to discern between visually similar species. Shell colour and pattern of the shell can help. However, this can be varied between individuals of the same species. Empty shells can also have a different appearance than those with the snail inside. Other useful features can include the direction and number of whorls, shell thickness, surface sheen and texture. 

Very little equipment is needed for identifying snails, but a hand lens can help for smaller specimens, particularly when counting whorls or looking at shell textures. Specimen pots or trays can help you to safely store species while you study them, and forceps are useful for collecting and moving smaller, more delicate species.   

Garden Snail (Cornu aspersum)

Garden Snail on a leaf in a garden.
Snail in our garden by Les Pounder, via flickr.

Distribution: Common throughout lowland Britain. 

What to look for: This is a well-known species that most people will have seen in their gardens or local green spaces. The garden snail has a thick shell, with a mottled brown, red, and yellow colouration. Its shell aperture is large and has a thickened white lip. It has around 4.5–5 whorls and its thick shell has a rough, wrinkled surface. The umbilicus, the depression or hole at the centre of shell whorls, often on the underside, is completely sealed by the lip. 

White-lipped Snail (Cepaea hortensis)

White-lipped snail on concrete by hedera.baltica.
White-lipped snail by hedera.baltica, via flickr.

Distribution: Widespread across Britain, but mainly found in coastal areas in Scotland. 

What to look for: This species has a glossy, smooth shell that is usually a yellow colour. However, individuals can be pink, brown or red. The number and presence of dark spiral bands can vary but there is no more than five. This species most often has an obvious white lip around the shell aperture.  

Brown-lipped Snail (Cepaea nemoralis) 

Brown-lipped snail travelling across a concrete pavement.
Brown-lipped snail by hedera.baltica, via flickr.

Distribution: Widespread across Britain apart from the northern parts of Scotland. 

What to look for: The colouration of this snail is widely variable and can be yellow, brown or pink. The presence of the banded patterning is also variable and they can have up to five bands across their shells. Their shells have between 4.5–5.5 whorls, with a semi-glossy surface. There is usually a dark rim to the lip of the shell aperture.   

Hairy Snail (Trochulus hispidus) 

Trochulus hispidus - Hairy Snail climbing up a branch.
Trochulus hispidus – Hairy Snail by Nikk, via flickr.

Distribution: Widespread throughout Britain. 

What to look for: This snail can vary in colour from cream to brown. It sometimes has a light band around the shell aperture. The shell is quite flat and densely covered in short hairs, which can be worn away over time. These hairs have been found to help the snail to adhere better to wet surfaces.  

Copse Snail (Arianta arbustorum) 

Heesterslak - Arianta arbustorum snail on concrete.
Heesterslak – Arianta arbustorum by Gertjan van Noord, via flickr.

Distribution: Widespread. 

What to look for: This species can grow up to 19mm. Its shell is a mottled brown with a thin band around the circumference, although its colour pattern can be highly variable. Its body is very dark and the shell aperture is a ‘C’ shape, often with a paler inside lip that can be bone-white. The shell has between 5–6 whorls and the umbilicus is a small crescent-shaped slit.  

Kentish Snail (Monacha cantiana) 

Kentish Snail (Monacha cantiana) on a green leaf.
Kentish Snail (Monacha cantiana) by Peter O’Connor, via flickr.

Distribution: Widespread across England, less common in Wales and Scotland. 

What to look for: This non-native species has a creamy shell with dark mottling. It often has a pale band around its circumference and a relatively small umbilicus. The body of the snail is a pale brown, with a darker skirting and sometimes darker tentacles.  

Striped Snail (Cernuella virgata) 

Snail at Walkley, Sheffield, crawling across stones.
Snail at Walkley, Sheffield by Tim Parkinson, via flickr.

Distribution: Widespread.  

What to look for: Also known as the vineyard snail, this snail has a pale shell, usually with dark spiral bands. The shell colouration and the number of markings are variable. It is an uncommon species, usually found in calcareous grassland, sand dunes and coastal grasslands.  

Pointed Snail (Cochlicella acuta) 

Pointed Snail attached to a tree.
Pointed Snail by Katja Schulz, via flickr.

Distribution: Found mainly in Wales, Ireland, and south and west England, it also occurs on some islands off of Scotland. 

What to look for: It has an elongated conical shell that tapers to a blunt tip. This shell varies in colour and markings but is usually a pale cream or off-white. It may have several bands of dark brown or black or be streaked with brown. 

Amber Snail (Succinea putris) 

Succinea putris, large Amber Snail, on the fold of a green leaf.
Succinea putris. Large Amber Snail by gailhampshire, via flickr.

Distribution: Widespread throughout England and Wales, less common in Scotland. 

What to look for: Between 15–22mm tall and 7–12mm wide, the shell of this species can range from very light amber to a darker orange-brown in colour. Its shell also has a very large final whorl. The body of this snail is a pale colour with two dark lines running along the top of its head, extending along its tentacles to its eyes. 

 

The NHBS Guide to UK Slug Identification

Slug is the common name for gastropod molluscs that have little to no shell. Descended from snails, they usually have a vestigial shell that is internalised, but some have either none at all or a very reduced one, such as the shelled slug (Testacella scutulum), which has a fingernail-like shell over its rear end. There are over 30 species of slug in the UK. 

As they do not have full-sized shells, they’re prone to desiccation, so many species are most active during and after wet weather and spend drier times hidden in damp places such as under man-made structures, tree bark, leaf litter and rocks. They play an essential role in the ecosystem, similarly to snails, by eating decaying matter such as plant material and fungi, aiding nutrient cycling.  

A small number of slugs are considered serious pests to agriculture and horticulture, eating foliage, fruits, and vegetables. This gave rise to the widespread use of toxic slug killing chemicals, which often impacted other non-target species. Recently, however, the use of iron phosphate baits has emerged, as they are less harmful to other wildlife.  

Black Slug (Arion ater) 

Arion ater. Large Black Slug, on grass.
Arion ater, Large Black Slug by gailhampshire, via flickr.

Distribution: Extremely common and widespread throughout Britain. 

What to look for: These species are large and vary widely in colour, including black, brown, grey, orange, reddish and green. It has long, coarse tubercles, the raised areas between the grooves on its skin, found on its side and back. Black slugs also have a pneumostome, a breathing hole, on the right side of their mantle, a protective structure of calcareous granules, through which they breathe.  

Common Garden slug (Arion distinctus) 

Arion distinctus on a rock.
Arion distinctus by Donald Hobern, via flickr.

Distribution: Widespread.  

What to look for: A dark or bluish-grey with dark lateral stripes and a pale yellow or orange sole (underside), with a characteristic yellow-orange mucus and no keel. This species also has tiny gold speckles on its tubercles, which are best seen through a hand lens. 

Netted slug (Deroceras reticulatum) 

Netted Slug - Deroceras reticulatum sliding down a leaf.
Netted Slug – Deroceras reticulatum by AJC1, via flickr.

Distribution: Widespread. 

What to look for: This may be the most common slug across the UK. It is a pale, off-white colour, with a keel at the tip of its tail and a mantle that is roughly half the length of its body. It has a chunky build and the tubercles are pale than the rest of its skin, giving it a netted appearance.  

Western Dusky slug (Arion subfuscus) 

Slug sliding across a leaf from left to right.
Slug by Rob Mitchell, via flickr.

Distribution: Widespread, but less common in East Anglia. 

What to look for: The dusky slug has various colour forms of yellow and brown. An important feature is the orange body mucus which stains on contact with your skin. It also usually has two dark lateral stripes and a pale sole with a fringe that blends in with the body.  

Hedgehog slug (Arion intermedius) 

Hedgehog Slug (Arion intermedius) crawling along some mud.
Hedgehog Slug (Arion intermedius) by Richard Ash, via flickr.

Distribution: Widespread but not common. 

What to look for: This is a small slug, at only 1522mm when extended, and is usually pale brown with a darker lateral stripe along its body to the mantle. Its sole is pale yellow and it has coarse tubercles that can contract to ragged points, giving it the reason for its name.  

Leopard slug (Limax maximus) 

Leopard slug on a marble stone.
Leopard slug by David J, via flickr.

Distribution: Widespread. 

What to look for: The appearance of the leopard slug is quite variable but it usually has a pale background and distinctive dark spots and markings that resemble leopard spots. The markings on its back are usually arranged in three longitudinal bands and it has a pale sole.   

Yellow slug (Limax flavus) 

Yellow Slug (Limax flavus) in a back garden.
Yellow Slug (Limax flavus) by Peter O’Connor, via flickr.

Distribution: Found mainly in England and Wales. 

What to look for: The yellow slug is usually a bright, lemon yellow with darker markings, with blue tentacles and a yellow line along the keel which extends from the tail to approximately halfway along its body. This central line can sometimes be broken into dashes.  

Green Cellar Slug / Irish Yellow Slug (Limacus maculatus) 

Irish Yellow Slug (Limacus maculatus).
Irish Yellow Slug (Limacus maculatus) with slug mites by Peter O’Connor aka anemoneprojectors, via Wikimedia Commons.

Distribution: Introduced population with a spreading range across the UK, less common in Scotland. 

What to look for: This species can vary from green to dull yellow with dark markings, grey tentacles and a colourless to orange slime. It can also occasionally have a yellow stripe at the tail end that doesn’t reach more than halfway along its body.  

 

Recommended Reading: 

Slugs of Britain and Ireland: Identification, Understanding and Control   Paperback | Feb 2014

This richly illustrated guide provides a comprehensive, picture-based identification charts and species accounts for 46 slug and semi-slug species known in Britain and Ireland.

 

Slugs and Snails Hardback | December 2016

In this long-awaited New Naturalist volume, Robert Cameron introduces gastropods, sharing the comprehensive natural history of slugs and snails of the British Isles specifically.

 

 

 

Opticron Hand Lens 23mm 10x Magnification 

Perfect for fieldwork, this general-purpose hand lens is ideal for observing insects.

 

180ml Collecting Pot

Sterile, screw-on sampling containers ideal for viewing insects in the field.

 

Book Review: Of Cockroaches and Crickets

***** An amusing and light read

Of all the insects that have a PR problem, cockroaches must rank very high. That, however, did not stop German entomologist, journalist, and filmmaker Frank Nischk from spending a year-long internship studying them. In this book, he regales the reader with stories of his time in the lab and the field studying first cockroaches and later crickets. A light and breezy read despite the serious undercurrent of biodiversity decline, Of Cockroaches and Crickets turned out to be an entertaining read.

This book was originally published in German in 2020 as Die fabelhafte Welt der fiesen Tiere by Ludwig Buchverlag and has been translated into English by Jane Billinghurst who frequently works with Greystone Books. Carl Safina contributes a short foreword that cracked me up and immediately set the tone. The book is effectively a memoir of Nischk’s early years studying for his undergraduate and doctorate degrees in the mid-nineties, told in 18 short chapters in two parts. His subsequent career pivot to documentary filmmaking only receives passing mention.

Cockroach photographed in Australia.
Cockroach by Patrick Kavanagh, via flickr.

Given Nischk’s concern about biodiversity decline, and his desire to communicate to a broad audience why insects are fascinating and important, there is an irony to his undergraduate internship. He spent a year in the lab of Martin Dambach studying the aggregation behaviour of the German cockroach, Blattella germanica. By day, large groups of them bed down on their own excrement, likely attracted by pheromones released by the faeces. The irony? Nischk’s internship was funded by biotechnology giant Bayer which was hoping to isolate the chemicals responsible for putting the cockroaches in sleep mode to develop a pheromone-based cockroach trap: “the exterminator’s holy grail” (p. 25).

For his subsequent doctoral studies, Nischk got his conservation priorities in order. Staying with Dambach, he turned to crickets and spent time in Ecuador recording their songs. Next to discovering species new to science, this is his entry into the fascinating field of soundscape ecology or ecoacoustics. A small cadre of ecologists has been recording soundscapes of natural habitats. Bernie Krause (not mentioned here) is one particularly well-known example. By comparing recordings made years or decades apart they have shown how natural soundscapes are changing and often disappearing due to human encroachment. Others are hoping to train software to analyze recordings and identify species by their calls. If scaled up, the dream is to have passive acoustic monitoring stations in biodiversity hotspots around the globe.

This backbone of his research is livened up with personal anecdotes and interesting asides. A friend’s call about a wasp infestation in her kitchen drawer is an excuse to introduce the 18th-century French entomologist Jean-Henri Fabre who was one of the first European naturalists to systematically collect and study butterflies, beetles, and wasps. Getting stung by a bullet ant in the rainforest of Ecuador leads to an aside about the late entomologist Justin O. Schmidt, the man who got stung for science (and wrote a fine book about it too). Tracking down a particularly loud cricket in Ecuador is the starting point for an unusual case where entomologists helped to defuse international political tensions between the USA and Cuba (this story has a surprising twist that I will not spoil here). A botched attempt to eradicate cockroaches that escape his experimental setup backfires most spectacularly, while fieldwork in the tropics is always fodder for amusing cultural misunderstandings and sober reflections. There is a nice mix here that never dwells on any one topic too long and makes for a book that is hard to put down.

Cricket on a leaf in a garden.
Cricket by Dean Morley, via flickr.

The third and final part is, perhaps surprisingly, comparatively the weakest of the book. In four chapters Nischk muses on the biodiversity crisis, particularly the still poorly understood decline of insects, and discusses examples of individuals and organisations who are creating and protecting wildlife habitat. Probably most interesting are the little-known grassroots initiatives in Ecuador that are undertaken by villagers and farmers turning to ecotourism. But is this really the answer? Or does it merely perpetuate the idea that nature can only be protected if it has monetary value? You will not find a critical or comprehensive analysis of wildlife conservation here. There is also an odd focus on projects in the USA, e.g. the High Line in New York, the Xerces Society, Joan Maloof’s Old-Growth Forest Network, and the Yellowstone to Yukon Conservation Initiative. I wonder if this was added for the English translation. There is no mention of e.g. the European Natura 2000 network of protected areas or E.O. Wilson’s bold call to protect half the planet, and only passing mention of the German environmental organisation NABU or the practice of rewilding. Putting aside such nitpicking, none of this takes away from his genuine concern about the ongoing loss of biodiversity nor from his conclusion that the key to protecting species is protecting their habitat.

Overall, Of Cockroaches and Crickets is an amusing and light read that I devoured in a day. Nischk offers a nicely balanced blend of interesting natural history, amusing personal stories, and captivating scientific research. Whether it is flies, wasps, or rats, we need more books that celebrate those species we all too readily dismiss as pests.


Of Cockroaches and Crickets book cover.Of Cockroaches and Crickets is available from our online bookstore.

Author interview with Phil Sterling and Mark Parsons: Field Guide to the Micro-moths of Great Britain and Ireland

Part of the popular Bloomsbury Wildlife Guides series, the first edition of the Field Guide to the Micro-moths of Great Britain and Ireland is credited with helping to demystify the formerly obscure and specialist world of micro-moths. Among a number of pioneering features, the book boasted artwork by acclaimed illustrator Richard Lewington showing the moths in their natural resting positions, rather than as pinned specimens, capturing the essence of each species and presenting them as they would appear in life when encountered in the field or settled within a moth trap. Coupled with accessible species accounts and the latest distribution maps, the guide helped to showcase the beauty and diversity of these miniature lepidopterans and provide a gentle introduction to their identification and habits for the general naturalist.

Eleven years on, the team behind the original guide have produced an extensively revised second edition, due to be published in December. Here the authors, Phil Sterling and Mark Parsons, kindly took the time to answer some of our questions about what we can expect from the new guide, and about micro-moth recording more generally.

Authors Mark Parsons (left) and Phil Sterling (right)

To begin with, could you give us a brief overview of what’s changed in the second edition?

There are three stand-out changes for recorders browsing this second edition. Firstly, we have included many more species, nearly 300 of them; the field guide now covers over 80% of native, naturalised or immigrant species recorded in Great Britain and Ireland. That’s 1,286 from a total of 1,576 species, and the book features over 1,000 of Richard Lewington’s illustrations.

This time we have included a full set of common (or vernacular) names for the micros which appear alongside the scientific names. We reviewed existing common names being used, made changes where we felt these were appropriate and helpful, and have done this for all 1,576 species.

The order in which the species are presented is now in line with the Agassiz, Beavan & Heckford (ABH) Checklist of the Lepidoptera of the British Isles. The ABH checklist was published in 2013, a year after our first edition, and it set out a radically different order based on our current understanding of the taxonomic relationship between species, genera and families. This should make it much easier for recorders to relate the order in the field guide to established recording systems and websites.

Box-tree Moth by Phil Sterling

You mention that the species coverage has been greatly expanded since the first edition. Where have these ‘new’ species come from?

There are a variety of reasons why we include more species. Most obvious are those we see today but did not 11 or more years ago. In 2007 when we started writing the first edition, Box-tree Moth hadn’t been recorded in Britain, yet it is now an abundant species across much of southern England and set to spread fast. Our warming climate is also leading to rapid increases in micro-moth species establishing and spreading, and we have included several scarcer immigrant species which do seem to becoming more regularly recorded.

We will never be up-to-date though. No sooner had the second edition gone to print than Echium Stilt Dialectica scalariella made a spectacular appearance in Britain in autumn 2023. It now appears to be resident at various sites from Kent to Devon, and possibly Suffolk, and has been found in huge numbers as a leaf-miner on Viper’s-bugloss, disfiguring the leaves. This species is listed in the field guide as British but not covered in detail.

We have taken a more systematic approach to inclusion of those that can be identified to species from their early stages, in particular leaf-miners in the Nepticulidae and Gracillariidae, and cases of the Coleophoridae. We have chosen a wide selection of these, illustrated by good quality photographs, enabling identification to species based on the early stage.

We have responded to helpful criticism that we should have been more comprehensive in our coverage of some of the more difficult families, including Gelechiidae and Depressariidae. There is better knowledge ‘out there’ today of how to separate species in the field, and we have been able include this information for some of the more challenging species within these groups.

There will be recorders who would have wished we included yet more species, maybe even all of them? But we have kept faithful to the objective of this being a field guide, for recorders to take it whenever they are out, by day and night. Most of the remaining 20% of micros require detailed examination under a microscope, extended time to rear the species from larva to adult to confirm identity, or are very rarely encountered species.

The bottom line is that we requested to Bloomsbury and they allowed us to write a bigger book than the first edition. We have packed it to our limit with more species and more information.

Mine of the Echium Stilt on Viper’s Bugloss by Mark Parsons.

For me (and I suspect many other people), the first edition of this guide was the catalyst to start paying more attention to micro-moths, having previously written them off as ‘too difficult’. How do you feel interest in micro-moths has changed in the 11 years since the original was published?

The first edition certainly achieved its aim of bringing micro-moths within comfortable reach of many recorders who had perhaps previously not considered them. Many species are actually straightforward to identify, but we think recorders were put off by the need to acquire several expensive texts to gain comprehensive coverage.

Moth recording generally continues to grow and it has wide public appeal. The National Moth Recording Scheme (NMRS), run by Butterfly Conservation, collates moth records across the four countries in Britain and Ireland and does not separate macro- and micro-moth species. We feel the distinction is fading, with recorders deciding which micro-moth families they’ll tackle in the coming year, rather than considering the prospect too daunting.

Leaf-mine of the Sorrel Dot Enteucha acetosae on Sheep’s Sorrel by Ben Smart.

Has that growth in interest in micro-moths helped you in your revisions to this guide, e.g. by helping to fill gaps in distribution maps and so on?

The first edition was primarily aimed at moth trappers, and we included lots of photographs of leaf-mines and larval cases to add interest. In the second edition we have deliberately chosen photographs of the early stages where identification to species level is possible based on a photograph and careful observation. We want to extend recorders’ interests beyond their moth traps to include searching for leaf-mines, cases and other signs in the field. Daytime pursuit of micro-moths can happen 365 days of the year, and there are plenty of species detectable in the middle of winter. The second edition benefits especially in this respect from photographs by Ben Smart, who has already published two small volumes on Micro-moth Field Tips, both giving month-by-month hints on how to find the early stages. Having said that, the second edition remains very much a book for moth trappers, and we have added illustrations of many more adult moths from a range of families.

Black-barred Fungus Moth Nemapogon clematella by Phil Sterling.

Could you tell us a little bit about the motivation and process behind your efforts to create the standardised list of common names in this guide?

Common names of micro-moths have never been adopted widely by recorders in Britain and Ireland. There is a long history to them, with a list first appearing in the early 17th century, and the most recent compilation was collated by Jim Wheeler in 2017. Despite the efforts, it is a ‘Marmite’ subject, with common names being used by some recorders yet remaining deeply unpopular with others. Much of the angst seems to centre on the inappropriateness of some names. All members of the Tineidae have been called ‘clothes moths’ yet this wonderfully diverse group contains only a very few species that will eat clothes, the majority are in fact detritivores and herbivores, with a range of specialists on fungi. Labelling them all as if they are clothes-munchers is just plain wrong at best, and gives moths a bad name at worst. Each fungal specialist is, not surprisingly, called a type of ‘fungus moth’, a simple revision based on its life history. There are other names that today appear to be meaningless, such as ‘tubic’, which gives recorders little to grasp when trying to remember them.

The introduction of a revised set of common names, in tandem with the increasing interest and our continued desire to promote micro-moth recording to all, seemed timely with the advent of the second edition. Very many common names remain as they are in the Wheeler list, or are slightly modified, and we have tried to make revisions that add relevance to the species, such as what it looks like, or where and how it lives. We hope that the names will make micros even more accessible by removing the perception that they somehow require more specialist knowledge to tackle.

Most recorders now use social media to communicate their finds, and it is undoubtedly easier to write an English common name than the scientific equivalent, and we have tried to keep the names as short as possible with this in mind. In summary, we hope that the revised list will be adopted in time and will encourage an inclusivity for new recorders put off by scientific names, and we trust we are putting micro-moths alongside almost all groups of British wildlife that now have common names.

Phil Sterling searching for larvae on Alderney by Peter Costen.

For any enthusiasts who have not yet looked beyond the more familiar macro-moths, what can you say about the rewards of taking the plunge into the world of micros?

Macro-moth recorders are delighted when they find something of great interest in their garden moth trap, it’s exciting, and it can happen to new and seasoned recorders. Imagine increasing your chances of this happening threefold! There are roughly twice as many micro-moth species as macros, plus their distribution is generally less well known, so the chances of finding ‘goodies’ is much enhanced. The advent of pocket-sized cameras and mobile phones with decent macro-photographic capability means that it is easier to share photos of micro-moths with others than it has ever been. Add to that the opportunity to record so many species from their early stages, and at any time of year, what are people waiting for?

Records of micro-moths are collated by the NMRS. By encouraging recording the amount of data collected will increase, along with our understanding of distributions and phenology. It won’t be long, we hope, until we can see national distribution maps for micros just like the macros.

Finally, are there any other projects you’re working on that you’d like to tell us about?

For us there is the continued quest to resolve undescribed life histories of micro-moths. Even some species seen regularly as an adult are not known as a larva, such as Dingy Knot-horn Hypochalcia ahenella. It probably feeds on herbs within its favoured habitats, sparsely vegetated stony or sandy ground. For what must be quite a large larva it is certainly secretive and has eluded us to date.

One mystery partly revealed in 2023 was the finding of a larva by Phil of Black-spotted Nest Moth Tenaga nigripunctella. The life history of this very rare tineid moth was completely unknown anywhere in the world and it was assumed that the larva might feed underground in rats’ nests. The adults have been appearing more regularly in west Dorset in the last few years, and a few were observed in a hole in a stone wall at Abbotsbury in 2023. In the wall was a dead Garden Snail Cornu aspersum, and guess what; deep within the shell a fully-fed larva of the moth that appeared to have been eating the blackened dead flesh of the snail. As far as we are aware this is the first example of a moth in western Europe found to be eating dead snails. There is so much to learn about the intriguing life histories of our more obscure micro-moths!


Field Guide to the Micro-moths of Great Britain and Ireland (Second Edition) is due to be published by Bloomsbury in December 2023 and is available from nhbs.com.

Book review: The Mind of a Bee

***** Fascinating and information dense
 Leon Vlieger, NHBS Catalogue Editor

It is tough being a social insect. When people are not trying to exterminate you, they might marvel at the collectives you form, but does anybody think much of you, the individual? Leave it to Lars Chittka, a professor in sensory and behavioural ecology, to change your views. The Mind of a Bee is a richly illustrated, information-dense book that explores a large body of scientific research, both old and new.

Chittka is very focused in his approach and The Mind of a Bee effectively summarises a large number of experimental studies in narrative form, with very few diversions. He cleverly avoids overheating your brain by having chapters flow logically into each other, but especially by dividing each chapter into short, headed sections. Each of these takes a particular question and discusses a few relevant studies in anywhere from one-half to three pages. Some of these are his own work but he ranges far and wide and includes both classic and recent research. The book is furthermore illustrated with numerous diagrams and photos that helpfully clarify experimental protocols and results. Honey bees are unsurprisingly the most intensively studied but Chittka discusses informative studies across a range of bee species and sometimes other insects as well. The book roughly covers three biological disciplines: sensory and neurobiology, ethology, and psychology.

Justifiably, the book opens with sensory biology. Before we understand what is in the mind of any organism, Chittka argues, we first need to understand the gateways, the sense organs, through which information from the outside world is filtered. These are shaped by both evolutionary history and daily life (i.e. what information matters on a day-to-day basis and what can be safely ignored). Chapter 2 deals with the historical research that showed that bees do have colour vision and furthermore can perceive ultraviolet (UV) light. Chapter 3 bundles together research on numerous other senses, including ones familiar (smell, taste, and hearing) and unfamiliar to us (perception of polarised light, Earth’s magnetic field, and electric fields). The antennae of bees, in particular, are marvels; Chittka likens them to a biological Swiss army knife, packing numerous different sense organs into two small appendages. Tightly connected to sensory biology is how this incoming information is processed in the brain, though Chittka postpones discussing neurobiology to chapter 9. He describes the discovery and function of different brain areas and highlights the work of Frederick Kenyon who would inspire the better-remembered Santiago Ramón y Cajal. Thanks to them, we now understand that brains consist of numerous specialised nerve cells. Though the bee brain is small, Chittka argues that size is a poor predictor of cognitive skills; it is the wiring of neurons that matters. Rather than be surprised that small-brained insects such as bees can do so many clever things, Chittka instead tickles the reader with the opposite question: “Why does any animal need as large a brain as a bee’s?” (p. 153).

What clever things do bees do, you ask? That is the subject of the preceding five chapters where Chittka surveys a large body of behavioural research. Honey bees are famous for their waggle dance by which they communicate the location of flowers but also, this was news to me, the location of potential nest sites when the swarm relocates. But Chittka discusses more, much more: how bees navigate space using landmarks, show a rudimentary form of counting, solve the travelling salesman problem, learn to extract nectar from complex flowers, learn when to exploit certain flowers (and when to ignore them), and learn new tricks by observing other bees. But what about instinct, something most behaviours were traditionally ascribed to? He has some insightful comments on this: “even the most elemental behavior routines need to be refined by learning: instinct provides little more than a rough template” (p. 50). What really made me fall off my chair is that bees have long been outsmarting researchers in choice experiments. Many behavioural experiments take the form of choice tests, where bees need to pick between two locations or objects that differ in e.g. colour or shape with one option containing a sugary solution as a reward. Bumblebees would simply be lazy and check out both options in random order. Until, that is, protocols were modified by adding a bitter-tasting solution to the wrong choice as a penalty.

The final two chapters explore bee psychology. One chapter shows how, in a hive full of bees, the members are not anonymous and interchangeable. Rather, they show individual differences in e.g. their preferred order in which to visit flowers during foraging or how fast they learn to solve problems. The final chapter makes the case that bees have a form of consciousness, though Chittka clarifies he is not arguing it is as rich and detailed as that of humans. That said, they show a slew of behaviours that scientists will label as evidence for consciousness when exhibited by bigger-brained vertebrates. Chittka is happy to play devil’s advocate: sure, theoretically, all the behaviours described in this book could be replicated by an unconscious algorithm. However, the required list of specific instructions is growing long and, increasingly, the more likely answer seems to be that bees possess “a consciousness-based general intelligence system” (p. 208).

As mentioned above, this book is focused. If you enjoy reading about the facts and the study system with minimal (autobiographical) diversions, Chittka has got you covered. The only digression he allows himself is to include biographical details of older generations of scientists. This includes inspiring tales such as Karl von Frisch who described the honey bee waggle dance and later barely escaped being dismissed from his post by the Nazis. And look out for repeat appearances of Charles Turner, a now largely forgotten African American scientist who published pioneering work despite having been denied a professorship based on his ethnicity. But there are also tragic stories such as Kenyon’s, who snapped under pressure of not securing a permanent job and was incarcerated in a lunatic asylum where he died more than 40 years later, alone and forgotten. Chittka includes occasional quotations from historical literature to show that “many seemingly contemporary ideas about the minds of bees had already been expressed, in some form, over a century ago” (p. 15).

The Mind of a Bee makes for fascinating reading, convincingly showing that bees are anything but little automatons. The tight structure and numerous illustrations make it accessible, though be prepared for an information-dense book.

Author interview with Tim Blackburn: The Jewel Box

Interwoven throughout with tales of his experiences moth trapping on his London rooftop and in the Devon countryside, The Jewel Box by Tim Blackburn introduces us to a range of ecological theories and explains some of the where, why and hows that anyone curious about the natural world might tend to ponder upon. Beautifully and engagingly written, it manages to be both an ode to both the moths themselves and the activity of moth trapping, as well as a wider ranging exploration of the relationships between humans, other species and habitats.

Professor Tim Blackburn is a scientist with thirty years of experience studying questions about the distribution, abundance and diversity of species in ecological assemblages. He is currently a Professor of Invasion Biology at University College London, where his work focuses on alien species. Before that, he was the Director of the Institute of Zoology, the research arm of the Zoological Society of London.

In this Q&A we chatted with Tim about The Jewel Box as well as about moths in the UK, the things we still have to learn about them, and the species that he’s hoping to see in the flesh.


What struck me most about your book was how you manage to write about complex ecological theories in a very accessible way, while at the same time conveying your very personal admiration and fascination for these insects. What was it that convinced you that this book in particular needed writing?

For a few years now I’d wanted to write a book that presented the natural world in the way that ecological scientists tend to think about it. There is a lot of very fine writing about nature, but most of it is more natural history than science, or is very much focused on a specific organism or location. While The Jewel Box does use moths to illuminate and illustrate the rules that we (ecological scientists) think underpin how nature works, it is very much about those rules, rather than the moths themselves. I’m very happy to hear that you thought I explained the science in an accessible way – that was my fundamental goal.

Within the UK, we have a rich history of recording and studying moths. Where do you think are the big gaps currently in the research? Are there things about moths that we still know little about?

There are still lots of gaps in our knowledge of UK moths – hardly surprising given that we have 2,500 or so species here. For some, we still don’t know their natural food plant. For others, we don’t know if they still exist here. In this latter regard, it seems incredible to me that we are still arguing about the scale of insect declines in the UK, and what the causes of those declines are. We think moth numbers have probably dropped by 30% since 1970, but that information is only available for the commoner species of larger moths, and may be biased in various ways. While we have a rich history of moth recording, and some good data for moth population changes, we could really do with more.

Why do you think it can be so addictive to observe, identify and list the species that we find, whether it’s birds, plants or moths?

I don’t know! For me, it’s pretty much a hard-wired instinct. My mother says I was pointing at birds before I could talk. I’ve been a birder all my life. More generally, there is something deeply satisfying about observing and identifying species – a series of puzzles to work out. Yet unlike most puzzles, the solution is not a product of the human mind, but something more profound. It’s the start of an exploration into understanding millions of years of evolution and ecology. I love a cryptic crossword, but identifying moths gives me so much more joy.

As someone who was trapping before, during and after the Covid restrictions, did you observe any significant differences in the numbers or species of moths that were attracted to your trap during the periods of lockdown?

I wouldn’t say I noticed obvious differences due to lockdowns, although the first lockdown period itself was a very productive one for my moth trap. That was because I was in the Devon countryside when lockdown happened, rather than my upstairs London flat. The countryside is so much better for moths (numbers of species and individuals) than the city, and that spring was notably warm and sunny, which the moths loved. The 2020 lockdown was my most intensive period of trapping to date.

We are generally well informed about planting wildflowers for pollinators such as bees and butterflies and providing food for our garden birds. But what can we do to encourage moths?

The same really! Butterflies are just showy, diurnal moths. Moths are as good pollinators as bees, and like flowers and pollen just as much. So anything you do for the butterflies and bees will likely help the moths too. You just won’t see the impact as obviously, because most of the moths are using your garden while you’re sleeping.

Are there any species that you’ve yet to trap but are on your mothing ‘bucket list’ so to speak?

In The Jewel Box, I spoke about dreaming of catching a Death’s-head Hawk-moth, one of the largest and most iconic British moth species. Last October, I opened my moth trap on Blakeney Point in Norfolk to see that that dream had come true. It’s a moment that will stay with me forever. Now, Oleander Hawk-moth– the species that inspired the book’s cover – would be the dream, albeit even less likely than catching the Death’s-head.

Finally, what’s in store for you next? Do you have plans for further books?

I’m mulling on the next book, but still enjoying all that’s new and interesting in my life as a result of publishing The Jewel Box. But watch this space…


The Jewel Box by Tim Blackburn was published by Weidenfeld & Nicolson in June 2023 and is available from nhbs.com.