Hoopoe – A blog by nhbs – Page 12 – The NHBS Conservation Hub

The NHBS Guide to UK Snail Identification

Snails are a common feature in our gardens and parks. You may have particularly noticed them if you have a vegetable or plant patch, as they feed on the leaves, flowers and fruits of many of our food plants. There are over 40,000 species of land snail, although only around 120 occur in Britain.

There are several useful features for identifying the correct species. The overall shape, in terms of the ratio of height to breadth, is important, as species can vary between a wide, round, flattened shape to tall and thin. The shape, colouration and thickness of the mouth of the shell can often be used to discern between visually similar species. Shell colour and pattern of the shell can help. However, this can be varied between individuals of the same species. Empty shells can also have a different appearance than those with the snail inside. Other useful features can include the direction and number of whorls, shell thickness, surface sheen and texture.

Very little equipment is needed for identifying snails, but a hand lens can help for smaller specimens, particularly when counting whorls or looking at shell textures. Specimen pots or trays can help you to safely store species while you study them, and forceps are useful for collecting and moving smaller, more delicate species.

Garden Snail (Cornu aspersum)

Garden Snail on a leaf in a garden. — Snail in our garden by Les Pounder, via flickr.

Distribution: Common throughout lowland Britain.

What to look for: This is a well-known species that most people will have seen in their gardens or local green spaces. The garden snail has a thick shell, with a mottled brown, red, and yellow colouration. Its shell aperture is large and has a thickened white lip. It has around 4.5–5 whorls and its thick shell has a rough, wrinkled surface. The umbilicus, the depression or hole at the centre of shell whorls, often on the underside, is completely sealed by the lip.

White-lipped Snail (Cepaea hortensis)

White-lipped snail on concrete by hedera.baltica. — White-lipped snail by hedera.baltica, via flickr.

Distribution: Widespread across Britain, but mainly found in coastal areas in Scotland.

What to look for: This species has a glossy, smooth shell that is usually a yellow colour. However, individuals can be pink, brown or red. The number and presence of dark spiral bands can vary but there is no more than five. This species most often has an obvious white lip around the shell aperture.

Brown-lipped Snail (Cepaea nemoralis)

Brown-lipped snail travelling across a concrete pavement. — Brown-lipped snail by hedera.baltica, via flickr.

Distribution: Widespread across Britain apart from the northern parts of Scotland.

What to look for: The colouration of this snail is widely variable and can be yellow, brown or pink. The presence of the banded patterning is also variable and they can have up to five bands across their shells. Their shells have between 4.5–5.5 whorls, with a semi-glossy surface. There is usually a dark rim to the lip of the shell aperture.

Hairy Snail (Trochulus hispidus)

Trochulus hispidus - Hairy Snail climbing up a branch. — Trochulus hispidus – Hairy Snail by Nikk, via flickr.

Distribution: Widespread throughout Britain.

What to look for: This snail can vary in colour from cream to brown. It sometimes has a light band around the shell aperture. The shell is quite flat and densely covered in short hairs, which can be worn away over time. These hairs have been found to help the snail to adhere better to wet surfaces.

Copse Snail (Arianta arbustorum)

Heesterslak - Arianta arbustorum snail on concrete. — Heesterslak – Arianta arbustorum by Gertjan van Noord, via flickr.

Distribution: Widespread.

What to look for: This species can grow up to 19mm. Its shell is a mottled brown with a thin band around the circumference, although its colour pattern can be highly variable. Its body is very dark and the shell aperture is a ‘C’ shape, often with a paler inside lip that can be bone-white. The shell has between 5–6 whorls and the umbilicus is a small crescent-shaped slit.

Kentish Snail (Monacha cantiana)

Distribution: Widespread across England, less common in Wales and Scotland.

What to look for: This non-native species has a creamy shell with dark mottling. It often has a pale band around its circumference and a relatively small umbilicus. The body of the snail is a pale brown, with a darker skirting and sometimes darker tentacles.

Striped Snail (Cernuella virgata)

Snail at Walkley, Sheffield, crawling across stones. — Snail at Walkley, Sheffield by Tim Parkinson, via flickr.

Distribution: Widespread.

What to look for: Also known as the vineyard snail, this snail has a pale shell, usually with dark spiral bands. The shell colouration and the number of markings are variable. It is an uncommon species, usually found in calcareous grassland, sand dunes and coastal grasslands.

Pointed Snail (Cochlicella acuta)

Pointed Snail attached to a tree. — Pointed Snail by Katja Schulz, via flickr.

Distribution: Found mainly in Wales, Ireland, and south and west England, it also occurs on some islands off of Scotland.

What to look for: It has an elongated conical shell that tapers to a blunt tip. This shell varies in colour and markings but is usually a pale cream or off-white. It may have several bands of dark brown or black or be streaked with brown.

Amber Snail (Succinea putris)

Succinea putris, large Amber Snail, on the fold of a green leaf. — Succinea putris. Large Amber Snail by gailhampshire, via flickr.

Distribution: Widespread throughout England and Wales, less common in Scotland.

What to look for: Between 15–22mm tall and 7–12mm wide, the shell of this species can range from very light amber to a darker orange-brown in colour. Its shell also has a very large final whorl. The body of this snail is a pale colour with two dark lines running along the top of its head, extending along its tentacles to its eyes.

The NHBS Guide to UK Slug Identification

Slug is the common name for gastropod molluscs that have little to no shell. Descended from snails, they usually have a vestigial shell that is internalised, but some have either none at all or a very reduced one, such as the shelled slug (Testacella scutulum), which has a fingernail-like shell over its rear end. There are over 30 species of slug in the UK.

As they do not have full-sized shells, they’re prone to desiccation, so many species are most active during and after wet weather and spend drier times hidden in damp places such as under man-made structures, tree bark, leaf litter and rocks. They play an essential role in the ecosystem, similarly to snails, by eating decaying matter such as plant material and fungi, aiding nutrient cycling.

A small number of slugs are considered serious pests to agriculture and horticulture, eating foliage, fruits, and vegetables. This gave rise to the widespread use of toxic slug killing chemicals, which often impacted other non-target species. Recently, however, the use of iron phosphate baits has emerged, as they are less harmful to other wildlife.

Black Slug (Arion ater)

Arion ater. Large Black Slug, on grass. — Arion ater, Large Black Slug by gailhampshire, via flickr.

Distribution: Extremely common and widespread throughout Britain.

What to look for: These species are large and vary widely in colour, including black, brown, grey, orange, reddish and green. It has long, coarse tubercles, the raised areas between the grooves on its skin, found on its side and back. Black slugs also have a pneumostome, a breathing hole, on the right side of their mantle, a protective structure of calcareous granules, through which they breathe.

Common Garden slug (Arion distinctus)

Arion distinctus on a rock. — Arion distinctus by Donald Hobern, via flickr.

Distribution: Widespread.

What to look for: A dark or bluish-grey with dark lateral stripes and a pale yellow or orange sole (underside), with a characteristic yellow-orange mucus and no keel. This species also has tiny gold speckles on its tubercles, which are best seen through a hand lens.

Netted slug (Deroceras reticulatum)

Distribution: Widespread.

What to look for: This may be the most common slug across the UK. It is a pale, off-white colour, with a keel at the tip of its tail and a mantle that is roughly half the length of its body. It has a chunky build and the tubercles are pale than the rest of its skin, giving it a netted appearance.

Western Dusky slug (Arion subfuscus)

Slug sliding across a leaf from left to right. — Slug by Rob Mitchell, via flickr.

Distribution: Widespread, but less common in East Anglia.

What to look for: The dusky slug has various colour forms of yellow and brown. An important feature is the orange body mucus which stains on contact with your skin. It also usually has two dark lateral stripes and a pale sole with a fringe that blends in with the body.

Hedgehog slug (Arion intermedius)

Distribution: Widespread but not common.

What to look for: This is a small slug, at only 15–22mm when extended, and is usually pale brown with a darker lateral stripe along its body to the mantle. Its sole is pale yellow and it has coarse tubercles that can contract to ragged points, giving it the reason for its name.

Leopard slug (Limax maximus)

Leopard slug on a marble stone. — Leopard slug by David J, via flickr.

Distribution: Widespread.

What to look for: The appearance of the leopard slug is quite variable but it usually has a pale background and distinctive dark spots and markings that resemble leopard spots. The markings on its back are usually arranged in three longitudinal bands and it has a pale sole.

Yellow slug (Limax flavus)

Distribution: Found mainly in England and Wales.

What to look for: The yellow slug is usually a bright, lemon yellow with darker markings, with blue tentacles and a yellow line along the keel which extends from the tail to approximately halfway along its body. This central line can sometimes be broken into dashes.

Green Cellar Slug / Irish Yellow Slug (Limacus maculatus)

Distribution: Introduced population with a spreading range across the UK, less common in Scotland.

What to look for: This species can vary from green to dull yellow with dark markings, grey tentacles and a colourless to orange slime. It can also occasionally have a yellow stripe at the tail end that doesn’t reach more than halfway along its body.

Recommended Reading:

Slugs of Britain and Ireland: Identification, Understanding and Control Paperback | Feb 2014

This richly illustrated guide provides a comprehensive, picture-based identification charts and species accounts for 46 slug and semi-slug species known in Britain and Ireland.

Slugs and Snails Hardback | December 2016

In this long-awaited New Naturalist volume, Robert Cameron introduces gastropods, sharing the comprehensive natural history of slugs and snails of the British Isles specifically.

Opticron Hand Lens 23mm 10x Magnification

Perfect for fieldwork, this general-purpose hand lens is ideal for observing insects.

180ml Collecting Pot

Sterile, screw-on sampling containers ideal for viewing insects in the field.

This week in biodiversity news – 4th March 2024

Rediscoveries and Recovery

The Norfolk hawker dragonfly is no longer extinct, with the species set to be taken off the British Red List of endangered species after a large-scale population surge. Since its extinction in the Cambridgeshire Fens in the 1890‘s, the Norfolk Hawker’s small population has been restricted to the Norfolk Broads. However, in recent years they have been found in Kent, Hertfordshire, Sussex, Dorset and Devon due to an increase in suitable habitats across Britain resulting from climate breakdown. This has been the driving force behind the species recovery, however, the gaps in the current population distribution need filling to ensure the Norfolk Hawker’s long-term survival.?

Norfolk Hawker Dragonfly perched on a plant in the sun. — Norfolk Hawker by Jo Garbutt, via flickr.

A breeding population of an incredibly rare species of turtle has been discovered in India. The Cantor’s Giant Softshell Turtle is native to south and south-east Asia and has been classed as critically endangered. The discovery of a breeding female on the banks of the Chandragiri River in Kerala, however, provides hope for this illusive species. The study was led by a group of international conservationists who worked alongside local communities to record the first documentation of a female nest, and subsequently rescue the eggs from flooding. The hatchings have since been released into the river, and the team are now working closely with the local community to set up a hatchery and nursery near the original nesting site.?

Climate

Since 2000, more than a third of Antarctica’s glacier anchors have reduced in size. These anchors, or “pinning points,” are important braking mechanisms that hold the glaciers in place and subsequently reduce the rate of ice detachment from the continent. New research conducted by the University of Edinburgh suggests that an increase in global sea temperatures is causing these points to thin at a much faster rate than previously estimated. Since 2000, 37% of pinning points have either shrunk or completely disappeared, and, as a result, the glacier’s braking force is weakening.?

Amazing shapes and deep blue tumbled down the huge glaciers that lined the shores. — Antarctica 162 – dramatic landscape by McKay Savage, via flickr.

The UK has had the wettest February on record, according to the Met Office. Between December 2023 and January 2024, the UK experienced 90% of the total expected winter rainfall. This increase in wet weather continued throughout February with some areas of the UK seeing two and a half times their normal rainfall for this month alone. These adverse weather conditions, a result of climate change, is having detrimental effects on farmers due to increased crop flooding, with some farms in Lincolnshire having been underwater since October.

Science and Environment

A 12mm long, transparent fish has been found to make a sound as loud as a pneumatic drill. Scientists in Berlin were studying the Danionella cerebrum in their lab when they began hearing strange clicking noises coming from the male fish in the tank. Upon investigation, they discovered that this miniscule species can make a sound of 108 decibels when measured at a distance of one metre away – this is roughly equivalent to the noise made by a bulldozer and is likely to be the volume at which other fish hear the sound. This noise is created by the contraction and release of the fish’s swim bladder, and it is one of the loudest noises ever discovered in a fish of this size.?

Ancient trees play an irreplaceable role in supporting forest life, according to experts. The results from a recent study in the Spanish Pyrenees show that ancient trees no longer exert energy on reproduction, and instead prioritise stress tolerance, durability and slower growth. It has also been revealed that, over time, sections of older trees die and decay, which allows them to host more diverse and greater quantities of different species than younger trees, as juveniles don’t possess the unique physical and physiological features to support them. Although the number of old growth trees continue to decline worldwide, researchers Munne-Bosch and Pasques hope their report will highlight the importance of our ancient trees’ ecological role.?

Old, twisted Silver Birch tree near Owler Tor in the Peak District. — Silver Birch by Steve Batch 61, via flickr.

Policy

The government are expected to announce a move to 100% badger culling under exceptional circumstances in the coming weeks. The current policy on badger culling aims to reduce badger numbers by 70% in each cull area, however the government may increase the target to 100% in certain circumstances, subject to consultation, as of January 2026. The Government had previously stated that the cull would end in 2025 and be replaced by increased badger vaccinations, however some fear that the new 100% cull policy may become standard practice instead.

Parliament has implemented the first EU nature law to restore 30% of the EU’s degraded ecosystems by 2030. This comes into effect after recent findings reveal that over 80% of all European habitats are in poor shape, and aims to ensure the restoration of 60% of these environments by 2040, and 90% by 2050. The EU has pledged that once these ecosystems are restored, they will ensure that the area does not significantly deteriorate once more in a bid to aid the long-term recovery of damaged natural areas.

Author Q&A with Illustrator Mike Langman

Mike Langman has been a full-time illustrator specialising in birds since 1992 and has published a total of 85 books, including Park and Garden Birds and the Guide to Ducks, Geese and Swans, with his work also featuring in many UK birdwatching magazines.

Mike worked for the RSPB at their headquarters in Bedfordshire for nine years after finishing his education at Middlesex Art College in 1983, and his illustrations have been published in most of the RSPB’s quarterly Nature’s Home magazines, on nearly every RSPB reserve, on identification cards, in murals in information centre, and in other outlets across Britain and Europe.

He has been an avid birdwatcher from the age of ten and particularly enjoys birdwatching around his local area in the South West, especially Berry Head in Devon, with his knowledge and expertise regularly sought by local organisations. Mike is also a voluntary art editor for Devon Birds, his local bird society, where he has previously held a number of roles in numerous years between 1994 and 2010.

Mike recently took the time out of his busy schedule to answer our questions about his career in illustration, which mediums he uses to create his drawings and more.

How did you get into illustrating nature?

A love of nature started with walks with my parents and visits to my grandparents – one set keen on birds and the other owned a farm where I could (more or less) have free range to explore. Drawing wildlife came from copying pictures I saw in books of wildlife that excited me, initially big cats but as I watched birds in gardens and countryside I would copy pictures from books of them too.

What are your favourite mediums for illustrating books?

Most of my published work is a mixture of watercolour for its freshness and clarity, but to speed up the process I (like many other published artists) use gouache too, this being opaque pale paint that can be put on top of darker watercolour to create feather edges and highlights etc. A watercolour purist uses the white of the paper to shine through pale areas!

Mike Langman's pencil and watercolour double page illustrations with annotations of a Warbler. — Warbler field sketch by Mike Langman.

How do you approach illustrating a bird you’ve perhaps never seen in real life?

I use whatever I can get my hands on, googling photos or videos (better for capturing character), but I still use skins (dead crudely stuffed birds) held in museums like Tring or even Exeter and Torquay.

How do you record a bird‘s behaviours, and have you seen any unusual behaviours from a common bird?

Yes, as a birder I’m always looking for not just unusual species but behaviours too –– an inquisitive mind leads to a better understanding of the species and I do illustrate behaviour in published work when required to.

What has been your favourite bird to illustrate that you keep coming back to?

That’s a tricky one, but I love seabirds and warblers. I suppose if I had to choose one it‘s the Firecrest for its character and colour – a truly stunning tiny bird. But, I must not forget our south Devon speciality, the Cirl Bunting, and, and I could go on!

Cirl Bunting field sketch in pencil and watercolour by Mike Langman. — Cirl Bunting field sketch by Mike Langman.

How has climate change altered your approach to projects?

I do a huge amount of illustration work from home but I do travel a bit too (although much more locally based than I used to be). At home I’m trying to be as carbon neutral as possible with solar panels, storage batteries, good house insulation and, last year, buying an electric car. When I do travel abroad I do as much wildlife watching as possible – visiting reserves and hiring guides which will help maintain some of the very important and often not so important biologically rich areas.

Do you see wildlife human conflict in your work?

Not so much in the publishing world, but as part of my secondary job as a wildlife guide here in Devon, running walks, tours and cruises, I have witnessed some terrible scenes. Dolphins caught in nets, seals with plastic rings and fishing gear snagged around their necks, jet skis harassing dolphins, fishermen throwing rocks at seals and even fishermen having ‘fun’ catching gulls with baited hooks. Away from the sea we have hedges cut by land owners at the wrong times of year, and housing developments on green land around Torbay in areas I used to watch wildlife...

What bird do you wish you had seen and why?

Pretty much every bird I haven’t seen – but I know that’s not a good thing, practically and in terms of harm to my carbon footprint, and inevitably it’s just not possible. So, I keep it local and look for anything that provides me with a challenge to find, identify and share with others. For more than 30 years I’ve wanted to find a Hume’s Warbler (a small and very rare eastern Siberian migrant) at my local lakes at Clennon Valley in Paignton (where I volunteer as part of the friends group). In December 2023 I did a double take when I heard one calling, ‘Che-wee, Che-wee’. Eventually after what seemed like an eternity I located and watched it, sketched it and shared the sightings immediately with others. The bird stayed around for over a week. I guess I need a new goal now…

What projects are you working on right now?

I’ve just finished the 3rd edition of the Helm Field Guide to the birds of the Middle East and painted its new cover too, which I’m very proud of. I’m also working on some wood etching images (my work completed on the computer!) for Greenspace designs for the Lower Otter Restoration Project.

A collection of books illustrated by Mike Langman can be found in our bookstore here.

Author Q&A with Hugh Warwick: Cull of the Wild

In Cull of the Wild, author Hugh Warwick investigates the ethical and practical challenges of managing invasive species to increase biodiversity. He explores the complicated history of species control over time, while also combining scientific theory and subtle humour, to explain the many issues conservationists face when trying to protect native species from their non-native rivals.

Hugh Warwick is an ecologist, conservationist and writer who specialises in the study of hedgehogs. He has published three books focusing on this species, however he recently expanded his field of study to include invasive species while writing Cull of the Wild. He has previously written for BBC Wildlife, New Scientist, The Guardian and The Daily Telegraph, is the spokesperson for the Hedgehog Preservation Society, runs courses on both hedgehog conservation, and lectures in creative writing.

Hugh recently took the time to talk to us about his book, including what inspired him to write Cull of the Wild, how he hopes his book will help future conservation methods, and more.

Firstly, what inspired you to write a book on the culling of invasive species?

The very first independent field work I did in 1986 – for the 3rd year project of my degree – took me to North Ronaldsay. My supervisor had a friend who was both the GP and the bird warden on this island, the most northerly in Orkney. He had noted that there seemed to be a correlation between the increase in number of imported hedgehogs (that was the postman’s fault!) and the decrease in breeding success of ground nesting birds – in particular the Arctic Terns. My work was to try and find out how many hedgehogs there were. This turned out to be around 500 – not the 10,000 the Daily Express had reported! The bird observatory organised an airlift to remove the hedgehogs after I had gone and I returned in 1991 to repeat the survey. Since then I have found the collision of ecological study with human concerns really fascinating.

This led me to become involved, rather inevitably, in the Uist hedgehog saga, where the RSPB and Nature Scotland were killing the hedgehogs for the very same reason they had been airlifted from North Ronaldsay. I quickly shifted from reporting on the story for BBC Radio 4, to joining the campaign against the cull, and then doing the research that helped stop it. But while all this was happening, I met a researcher from New Zealand who was telling me that they were killing hedgehogs over there (in the 1860s we sent a load of hogs over to help the colonialists feel more at home) I could see no reason not to kill those hedgehogs.

That apparent contradiction has been spinning in my head ever since and was the springboard for this book.

Close up photograph of a New Zealand Hedgehog walking through long grass. — New Zealand: Hedgehog, by Eli Duke via flickr.

How did you find researching such differing opinions on the subject, and have your preconceived opinions changed over the course of writing this book?

I found this book so very different from all the others I have written. Previous outings have always been with people who are just really pleased to talk – to share their enthusiasm for Beavers or Water Voles or owls. This was the first time where I felt like the initial communication was almost like a job interview – assessing whether I was suitable to talk to. Some people simply refused to talk, others were cagey. I guess I was quite naïve!

The journey of the book is basically one where my head and my heart are in constant debate. I lay out my ‘bunny-hugging’ prejudices at the start. I suppose I was hoping to find that all instances where animals were being killed could be dealt with in other ways.

One of the biggest lessons I learnt, though, was about how people, even ones with whom you have fundamental differences of opinion, share so much common ground. But you will never see that common ground if you charge in at the difference! I love that one of my potentially conflicting interviews ended up with the biggest argument being over who recorded the best version of Sibelius’ 5th Symphony!

In Chapter 1, you discussed that animals are shown to be continually developing to become more like humans – problem solving, making and using tools etc. Do you think this gradual form of humanisation may lead to more equal ‘rights’ between animal species and humans in relation to killing for conservation?

I would not look at this as humanisation – that is slipping back into the human exceptionalism that got us into this mess. Currently the degrees of cruelty we deliver to wildlife is arbitrary, based in large part on language and on the names we give them, even different members of the same species.

We need to recognise that these animals being killed experience pain, joy, fear and hope. If the realisation that different species have the capacity to do human-like things is enough to get this recognised, then that is some progress.

I have been gently eased away from the concept of ‘rights’ for animals and towards a desire to see cruelty minimised. Rights are important and I see their value – but here, I feel the best outcomes will emerge from a utilitarian desire to maximise good and minimise bad. Though that requires we have the idea that these animals, most of whom are simply in the wrong place at the wrong time, are individuals and can suffer.

A close up of a Red Squirrel stood on its hind legs on a rock eating a nut it's holding in it's front paws. — Red Squirrel, by Caroline Legg via flickr.

You must have visited many places and spoken to many different people about different species while you were writing this book. Were there any particularly memorable moments or experiences that that have stayed with you?

I did not get to some of the places in this book – time, Covid, money and a desire to minimise flying all conspired to keep the exotic interviews on Zoom. But the adventures I did have were all special.

Heading out along the coastline of South Ronaldsay, Orkney, with Spud – the most handsome labrador I have ever met – was magical. The wind blew, the surf crumped in from the Atlantic, and Spud, well, he followed his nose and we followed him as he tracked down evidence of Stoats.

The Stoat story is so important – it is about speed and trying catch this problem early – they were only found in 2011. But more than the practical, it is also about the buy-in of the local population. If the public are not onside, the conservationists can give up now. So it is about education and communication as much as it is about deciding which traps to use and where to site them.

A close up of a stoat climbing over a dark, wet, mossy log. — Stoat (Mustela erminsea) by big-ashb, via flickr.

In what way do you hope that Cull of the Wild will help future animal conservation efforts?

The overarching message I hope people get from Cull of the Wild is that ecology is both fascinating and VERY challenging when it is part of a conflict. Ecology is not given the level of respect that it should receive. It is always sidelined – economics and politics hog the headlines – yet both of those are but subsets of the wider ecosystem. Without a healthy ecosystem we are on a downward slope to disaster. So, I hope that people will read this book, maybe because they are outraged at the killing, and will then come away realising that ecology is very complex and should be treated with great respect.

I also hope that this book will convince people who hold very firm opinions – opinions that are often amplified by the bubbles in which we tend to linger – that people with differing views may well share many of the same values as themselves. And that to begin discussion at the common ground is the foundation of progress. Argument should not be about winning, but about making things better. Remember, it is quite possible that some of the things you know are wrong. A friend of mine turned up at a conference recently with a t-shirt saying ‘Don’t believe everything you think’. We should be humble enough to recognise we might be wrong.

Are you currently working on any other projects that you can tell us about?

As I write this, the lovely people at the publishers Graffeg are hoping I will stop promoting Cull of the Wild and finish the two books I owe them: one on bats and the other on nocturnal nature. I have another narrative nonfiction brewing – about as different to this book as imaginable. Something which will hopefully bring laughter into our love of nature.

Cull of the Wild is available to order from our online bookstore.

Book Review: Of Cockroaches and Crickets

***** An amusing and light read

Of all the insects that have a PR problem, cockroaches must rank very high. That, however, did not stop German entomologist, journalist, and filmmaker Frank Nischk from spending a year-long internship studying them. In this book, he regales the reader with stories of his time in the lab and the field studying first cockroaches and later crickets. A light and breezy read despite the serious undercurrent of biodiversity decline, Of Cockroaches and Crickets turned out to be an entertaining read.

This book was originally published in German in 2020 as Die fabelhafte Welt der fiesen Tiere by Ludwig Buchverlag and has been translated into English by Jane Billinghurst who frequently works with Greystone Books. Carl Safina contributes a short foreword that cracked me up and immediately set the tone. The book is effectively a memoir of Nischk’s early years studying for his undergraduate and doctorate degrees in the mid-nineties, told in 18 short chapters in two parts. His subsequent career pivot to documentary filmmaking only receives passing mention.

Cockroach photographed in Australia. — Cockroach by Patrick Kavanagh, via flickr.

Given Nischk’s concern about biodiversity decline, and his desire to communicate to a broad audience why insects are fascinating and important, there is an irony to his undergraduate internship. He spent a year in the lab of Martin Dambach studying the aggregation behaviour of the German cockroach, Blattella germanica. By day, large groups of them bed down on their own excrement, likely attracted by pheromones released by the faeces. The irony? Nischk’s internship was funded by biotechnology giant Bayer which was hoping to isolate the chemicals responsible for putting the cockroaches in sleep mode to develop a pheromone-based cockroach trap: “the exterminator’s holy grail” (p. 25).

For his subsequent doctoral studies, Nischk got his conservation priorities in order. Staying with Dambach, he turned to crickets and spent time in Ecuador recording their songs. Next to discovering species new to science, this is his entry into the fascinating field of soundscape ecology or ecoacoustics. A small cadre of ecologists has been recording soundscapes of natural habitats. Bernie Krause (not mentioned here) is one particularly well-known example. By comparing recordings made years or decades apart they have shown how natural soundscapes are changing and often disappearing due to human encroachment. Others are hoping to train software to analyze recordings and identify species by their calls. If scaled up, the dream is to have passive acoustic monitoring stations in biodiversity hotspots around the globe.

This backbone of his research is livened up with personal anecdotes and interesting asides. A friend’s call about a wasp infestation in her kitchen drawer is an excuse to introduce the 18th-century French entomologist Jean-Henri Fabre who was one of the first European naturalists to systematically collect and study butterflies, beetles, and wasps. Getting stung by a bullet ant in the rainforest of Ecuador leads to an aside about the late entomologist Justin O. Schmidt, the man who got stung for science (and wrote a fine book about it too). Tracking down a particularly loud cricket in Ecuador is the starting point for an unusual case where entomologists helped to defuse international political tensions between the USA and Cuba (this story has a surprising twist that I will not spoil here). A botched attempt to eradicate cockroaches that escape his experimental setup backfires most spectacularly, while fieldwork in the tropics is always fodder for amusing cultural misunderstandings and sober reflections. There is a nice mix here that never dwells on any one topic too long and makes for a book that is hard to put down.

Cricket on a leaf in a garden. — Cricket by Dean Morley, via flickr.

The third and final part is, perhaps surprisingly, comparatively the weakest of the book. In four chapters Nischk muses on the biodiversity crisis, particularly the still poorly understood decline of insects, and discusses examples of individuals and organisations who are creating and protecting wildlife habitat. Probably most interesting are the little-known grassroots initiatives in Ecuador that are undertaken by villagers and farmers turning to ecotourism. But is this really the answer? Or does it merely perpetuate the idea that nature can only be protected if it has monetary value? You will not find a critical or comprehensive analysis of wildlife conservation here. There is also an odd focus on projects in the USA, e.g. the High Line in New York, the Xerces Society, Joan Maloof’s Old-Growth Forest Network, and the Yellowstone to Yukon Conservation Initiative. I wonder if this was added for the English translation. There is no mention of e.g. the European Natura 2000 network of protected areas or E.O. Wilson’s bold call to protect half the planet, and only passing mention of the German environmental organisation NABU or the practice of rewilding. Putting aside such nitpicking, none of this takes away from his genuine concern about the ongoing loss of biodiversity nor from his conclusion that the key to protecting species is protecting their habitat.

Overall, Of Cockroaches and Crickets is an amusing and light read that I devoured in a day. Nischk offers a nicely balanced blend of interesting natural history, amusing personal stories, and captivating scientific research. Whether it is flies, wasps, or rats, we need more books that celebrate those species we all too readily dismiss as pests.

Of Cockroaches and Crickets is available from our online bookstore.

Author Q&A with Chantal Lyons: Groundbreakers

Big, messy and mysterious – crossing paths with a Wild Boar can conjure fear and joy in equal measure. In Groundbreakers, Chantal Lyons gets up close and personal with this complex and intelligent species in the Forest of Dean, and investigates the people across Britain and beyond who celebrate the presence of these animals – or want them gone. From Toulouse and Barcelona where they are growing in number and boldness, to the woods of Kent and Sussex where they are fading away, to Inverness-shire where rewilders welcome them, join Chantal on a journey of discovery as she reveals what it might take for us to coexist with the magnificent Wild Boar.

Chantal Lyons is a naturalist, writer and science communicator. Having grown up in the tidy countryside of Kent, her encounters with the growing rewilding movement opened her eyes to the potential for restoring nature in Britain, and inspired her to study the relations between people and Wild Boar in the Forest of Dean. She currently lives in Cheltenham, never too far from the boar.

Chantal recently took time out of her busy schedule to tell us about her first experience with a Wild Boar, her hopes and fears for the future of their populations in Britain, and more.

First of all, could you tell us a little bit about yourself and what led you to writing a book about Wild Boar in Britain?

Up until my mid-twenties I spent all my writing time churning out unpublishable fantasy and sci-fi novels, alongside a zig-zagging career in the charity sector. The journey to Groundbreakers was perhaps a ponderous one – it started when I went to the Forest of Dean in summer 2014 to research a Masters dissertation in environmental social science. The Forest of Dean was, and still is, one of the few places in the country where Wild Boar have returned since their extirpation from Britain around 700 years ago. I wanted to find out from local residents what it was like to suddenly find yourself living alongside a big, wild, and utterly unfamiliar creature. What I discovered from those interviews astounded me, and I was sure that some established nature writer would soon publish a book about the return of boar to Britain and what this meant for us. No one did. After seven years, I decided to send off a proposal to a publisher who I’d encountered on Twitter, and it snowballed from there.

Could you tell us about your first real life experience with a Wild Boar?

It was a long time in coming! While I was in the Forest of Dean doing the Masters research, I used every spare moment to explore the woods on my own, often following locals’ leads. It always came to nothing. The boar seem to be like cats – you seem to be more likely to meet them the less you want to. But eventually, at the tail end of summer, I went to a spot where I’d heard from someone that a sounder (a family group of boar, which is always led by a matriarch) had been foraging each evening. I heard them softly calling to each other, then there was a rustling in the bracken, and an adult stepped out onto the path to get a look at me. We stared at each other for about two seconds, she gave a belching alarm call, and then she vanished with her family.

Wild boar and 7 piglets lead down on a rocky bank in the Yorkshire Dales. — Wild boar at Bolton Castle in Yorkshire Dales National Park, by HarshLight via flickr.

I appreciated the ways in which you speak about rewilding, in particular the importance of nuance and accepting the unknown. Do you think that uncertainties about the best way to undergo reintroduction projects and their expected outcomes are a significant hindrance to their implementation?

I have heard many a time from rewilding practitioners that the inherent uncertainties in rewilding can make it especially challenging to gain funding, given that projects aimed at restoring nature have traditionally been expected to be able to set clear targets and end-goals. But beyond practicalities like money, I think the bigger challenge is the attitude that reintroducing species for rewilding purposes is too big an unknown, and therefore too big a risk. Of course we should aim to conduct as much scientific research and gather as much knowledge as possible, but we are running out of time to reverse the haemorrhaging of our biodiversity in Britain and globally. To me, it beggars belief that we seem quite happy to continue doing all kinds of things that we know are massively damaging the environment, but reintroducing species? Why, that’s a step too far!

Close up image of two wild boar walking through mud with their snouts and feet covered in wet mud. — Wild boar, by Marieke IJsendoorn-Kuijpers, via flickr.

One part of the book that particularly resonated with me was in ‘The Risks of Being Alive’ where you speak about how, for most modern humans, ‘nothing matters more in life than eliminating all risk to it – even at the cost of happiness’. What do you think it would take for humans to accept more wildness into their lives?

Maybe it’s a cliché to say this, but I think it hugely helps to engage children with nature from a young age, and to continue doing so as they grow up. That of course means ensuring that everyone can access nature.

Beyond that, the question of how to convince people to accept more wildness is an incredibly tricky one, especially in a place like Britain which suffers so badly from ‘ecological tidiness disorder’ (as Benedict Macdonald puts it in Rebirding). The problem is that most of us have not made the connection between our tidy sterilised surroundings and the loss of nature. There is a lack of understanding that the majestically bare plains of Dartmoor or our rolling green fields mean an absence of life; and that you need all manner of species, including big ones, to ensure healthy ecosystems. I didn’t realise this for years. But once you know, you can’t stop wanting to bang the drum.

What are your biggest hopes and worst fears for the future of Wild Boar populations in Britain?

I want the planned, country-wide reintroduction of Wild Boar to become accepted both politically and societally. In the meantime, I hope that the few boar we do have are allowed to thrive. That means more oversight of people carrying out legal shooting of them on their land, and – in the case of the Forest of Dean population – better censusing methods to ensure they are not over-culled (though some culling will always be needed, as the boar currently lack other predators).

What am I most afraid of? That very soon, a disease called African Swine Fever (ASF) which burns through pig populations like wildfire will make its way into Britain. Since Brexit, border controls on pork imports have been so lax that experts think it’s only a matter of time before ASF reaches us. Forestry England is primed to wipe out every last boar in the Forest of Dean if the disease is ever detected in the population. And once they’re gone, it seems very unlikely that they would ever be reintroduced (legally or illegally) again. We will miss a miraculous chance to kickstart the landscape-scale restoration of nature in Britain.

Wild Boar piglets running across the road in a national park park with the sun out. — Wild Boar Sus Scrofa, by Björn, via flickr.

Finally, what is occupying your time at the moment? Are there plans for further books in the pipeline?

I work full-time as a science communicator, which does narrow my research and writing windows! But I am currently working on a proposal for book number two. It’s intended to be something of an evolution of Groundbreakers, picking up a thread that often emerged during my research interviews with people, but which I couldn’t possibly have fitted into this first book…

Groundbreakers is available from our online bookstore.

Author Q&A with Robert Wolton: Hedges

In Hedges, Robert Wolton brings together decades of research and personal experiences from his farm in Devon to explore the ecology, biology, nature conservation and wider environmental values of the hedges in the British Isles. Containing over 300 photographs and figures, this latest addition to the British Wildlife Collection offers a detailed commentary on hedges and their importance in our landscape.

Robert is an ecological consultant and writer specialising in the management of farmland and associated habitats for wildlife. He is a former hedgerow specialist for Natural England, the founder, chair, editor and lead author of the Devon Hedge Group, has been involved in Hedgelink since it began, and has written a number of reports and articles specialising in hedges.

Firstly, can you tell us a little bit about yourself and how you came to be contributing to the British Wildlife Collection with a book on hedges?

I’ve had a life-long interest in natural history, even as a schoolboy I was a very keen birdwatcher. Later, at university, I trained as a zoologist with a strong interest in mammals, although subsequently I have been more involved with insects, especially moths and flies. It was perhaps inevitable that I should pursue a career in nature conservation. My passion for hedges was awakened when we bought a small farm in the heart of Devon, my wife, Paula, looking after the cattle and sheep while I went to the office. Initially it was the flower-filled meadows that drew me to the land, but I soon realised that the many thick hedges, full of different trees and shrubs, were glorious and just as special, particularly when I discovered that those small spherical nests I kept finding were made by Hazel Dormice. At that time, 30 years ago, hedges were very under-appreciated in the nature conservation world – there was a gap waiting to be filled and I was able to persuade my bosses in English Nature to allow me to become a part-time national hedge specialist, a role I continued to fill after the organisation morphed into Natural England. Partial retirement gave me the opportunity to write a book on my beloved hedges. I’d always dreamt of having a volume in the British Wildlife Collection, much admiring the series, so when Bloomsbury offered me the chance I jumped at it.

Robert Wolton, author of Hedges, photographed stood by a large pile of thin trees being used to construct a man-made hedge. — Robert Wolton making a hedge.

I tend to think of hedges as being man-made. But is there such a thing as a natural hedge? And if so, how do these come about?

Most hedges in Britain and Ireland are indeed man-made. Some, though, have grown up naturally along fence lines and ditches – these are termed spontaneous hedges and I think they are becoming more frequent, especially along the sides of roads and railway lines. Trees and shrubs, their seeds carried by wind, birds and mammals, can colonise strips of rough grassland remarkably quickly, often protected to begin with by brambles. It does not take many years before there’s at least a proto-hedge present, and after a decade or two it may be difficult to tell it was not planted. Another way hedges have come into being is through strips of woodland being left when land is cleared for agriculture. These are called ghost hedges. Their origin is often given away by the presence of unexpectedly high numbers of trees and herbs characteristic of ancient woodland because they have poor dispersal abilities.

I particularly enjoyed the chapter in your book on the origins and history of hedges. Do you think that the study of hedges can give us an insight into the natural and social history of our country?

Without a doubt. Throughout our countryside, away from the open moors and fens, the pattern of fields, as defined by hedges and sometimes drystone walls, allows the history of the landscape to be read, often going back centuries, even sometimes millennia. We are so fortunate in these islands still to have this landscape continuity – it has been lost over much of continental Europe. In places like Dartmoor, which I can see from our farm, layer upon layer of history can be unpicked through studying the networks of field boundaries, most of which are banked hedges. Some can be traced back to the Bronze Age, 3,500 years ago. This may be exceptional, but even so, most of the hedges in Britain, and many in Ireland, probably date back to Medieval times. They are a hugely important part of our cultural heritage. We love our hedges. This is evident not just in the countryside, but across our villages, towns and cities, in our gardens and parks. Hedge topiary is, after all, a national pastime!

Farmland hedge leading towards a forest at the end of a farm track along the right-hand side of a field. — Hedge, by Damien Walmsley via Flickr.

As you describe early on in the book, there are many different types of hedge, from those that consist of just a single species to very diverse multi-species ones, even ones that have been allowed to mature into lines of trees. Is there a type of hedge that is best for the surrounding wildlife and environment and that we should be trying to replicate or maintain as much as possible?

If you put me on the spot, I’ll answer this question by saying that thick, dense, bushy hedges are the best for wildlife, preferably with margins full of tussocky grasses and wildflowers. But really we should be thinking about what networks of hedges look like, because there’s no such thing as a perfect hedge. Different birds, mammals and insects like different conditions, and in any case you can’t keep a hedge in the same state for ever, however carefully you manage it. Basically, the trees and shrubs are always trying to reach maturity, and as they do so gaps develops beneath their canopies and between them. That’s when laying or coppicing are needed, to rejuvenate the hedge and make it more dense and bushy. A lot more research needs to be done on this, but probably, from a wildlife point of view, at least half of all the hedges in a network, say that covering a decent-sized farm, should be in this condition. On the other hand, from a climate perspective, where we need to capture as much carbon as quickly as possible, tall hedges with many mature trees are best. You can see there are tensions here, all part of the challenge of managing hedges well. Who said it was easy?

A dusty track running in a straight line with hedges on both sides and tall, narrow, straight trees behind the left hand hedge. — Into the Distance, by Dave S via Flickr.

As both a farmer and an ecologist, I’m sure you are more attuned than many to the conflicting needs of making a living from the land and managing hedges for the benefit of wildlife and conservation. Do you think financial incentives are the only way to encourage landowners and farmers to both plant more and maintain existing hedges?

Financial incentives like government grants will always be important to landowners and farmers because good hedges benefit society at large just as much as those who own and manage them. Things like plentiful wildlife, carbon capture, reduced risk of homes flooding and beautiful landscapes rarely bring in any income to offset costs, let alone profit – it is right that they are supported from the public purse.

Still, hedges can be of direct financial value to farmers through serving as living fences, preventing the loss of soil or providing logs and wood chips for heating. They can also increase crop yields through boosting numbers of pollinators and the predators of pests. To some extent, these direct benefits to farm businesses have been forgotten in recent decades in the drive for increased food production regardless of environmental cost, but they are now being appreciated much more as new ways of working the land, such as regenerative farming, catch on.

And we should not overlook the fact that more and more landowners and farmers are prepared to bear at least some of the costs of good hedge management simply because they gain huge satisfaction from healthy hedges and all the wildlife they contain. The pleasure of seeing a covey of Partridges or a charm of Goldfinches, or hearing the purring song of the Turtle Dove, cannot be priced.

A narrow, windy track going through a high sided hedge into the distance in a circle. — by Oli Haines.

Finally, how did you find the experience of writing this book, and will there be other publications from you on the horizon?

This is my first ‘big’ book, and I was apprehensive to say the least when I started writing it, in 2022. But with a lot of encouragement from my wife and friends I soon got into the swing of things. Challenging for sure but personally most rewarding – exploring new facets, checking information and trying to find the best way to pass on my enthusiasm for the subject. Above all, it felt good to share knowledge collected over many years. Bloomsbury’s support was invaluable, there’s no way I could have self-published. As to whether there are more books in me, I’m not sure. Perhaps one on hedges in gardens? There again, I have a passion for wet woodland, another habitat that’s been much neglected. It’s all too soon to say.

Hedges is available to order from our bookstore.

This week in biodiversity news – 19th February 2024

Climate Crisis

A new study reveals the Amazon rainforest’s worst drought on record in over half a century, with human-caused climate change acting as the driving force behind these weather extremes. The Amazon rainforest plays an important role in the removal of CO₂ from the atmosphere, and accounts for around 10% of the world’s species. However, scientists suggest the Amazon could soon reach a “tipping point”, with droughts such as these expected every 13 years if global temperatures increase by a further 2°C.

Conservation

Black Rhinos have returned to Kenya’s Loisaba Conservancy as species recovers following local extinction 50 years ago. This translocation project, the result of a collaboration between Loisaba Conservancy and Kenya Wildlife Service, was prompted by the news that Kenya’s 16 Black Rhino sanctuaries are rapidly running out of space due to a boost in population numbers. Twenty-one of these critically endangered Black Rhinos were released into a specially created fenced sanctuary that covers nearly half of Loisaba’s total area, marking an important moment in the long-term future of Black Rhino populations in Kenya.

Black Rhinoceros stood in a plain full of grass. — Black Rhinoceros, by Andy Morffew via Flickr.

Hedgehog populations are on the rise according to BBC Gardeners’ World survey. Previous reports have found that since 2000, the number of hedgehogs has fallen by 30%-75% depending on the area due to increased habitat loss, fragmentation, pesticides, and a reduction in food sources. However, the annual hedgehog survey highlighted a 33% increase in sightings, up from 31% in 2022, which has been aided by the instillation of ‘hedgehog highways,’ and nationwide rewilding efforts. CEO of the British Hedgehog Preservation Society, Fay Vass, stated that urban populations are still lower than they should be, however these results provide cautious optimism for the future of hedgehogs.

Environment

Hailed as one of the world’s most ambitious environmental policies, Biodiversity Net Gain came into effect in England on the 12th February 2024. It is now mandatory for all new developments in England to deliver at least a 10% net gain in biodiversity or habitat on all new sites. In light of the government target of building 300,000 new homes a year by the mid-2020s, it is hoped that the scheme will minimise or mitigate the environmental effects of these new developments.

Iceland may be entering a new volcanic era, with increasing volcanic activity reported across the island. The most recent short-lived eruption on the Reykjanes peninsular marks the third since December 2023, and the sixth since 2021. Researchers think this wave of volcanic activity may be following a pattern that could last for decades, or even centuries. Scientists are trying to work out how to predict future eruptions by tracking ground inflations as magma rises, from which they can estimate when the magma will break the surface.

Litli-Hrútur – Volcanic Eruption in Iceland July 2023, by Anthony Quaintano via Flickr.

Science and discoveries

The latest strain of bird flu has decreased Great Skua populations by 75% and Northern Gannet populations by 25% in the past 2 years alone, says the RSPB and British Trust for Ornithology (BTO). According to a recent report, the H5N1 strain has been noted as one of the biggest conservation threats facing numerous seabird species across both UK and Europe. The latest Seabird Count survey, published in November 2023 – which did not take into account the effects of bird flu – found that almost 62% of seabird populations were already in decline across the UK due to other factors such as fishing and offshore wind developments.

The most vulnerable migratory species are at risk due to increased human activity, with one in five threatened with extinction according to the UN. Despite the signing of the Convention on the Conservation of Migratory Species of Wild Animals over 40 years ago, researchers suggest that more focus has been placed on preventing illegal animal trading rather than upholding the law on illegal killing. The report recommends that countries need to establish key biodiversity areas on the migration pathways of endangered species, reduce infrastructure obstructing migration routes, create corridors of protected land, and restore 30% of degraded marine and land areas.

Sandwich Terns stood on a beach in a colony looking out to sea. — Sandwich Terns, by Gary Leavens via Flickr.

British Wildlife Collection: interview with series editor Katy Roper

The British Wildlife Collection is a series of beautifully presented books on all aspects of British natural history. Since the series began in 2012, it has covered such diverse topics as mushrooms, meadows and mountain flowers, and books have been written by some of Britain’s finest writers and experts in their field. Filled with beautiful images, these wide-ranging and well-researched titles are a joy for any naturalist who is passionate about British wildlife and landscapes.

Katy Roper is a Senior Commissioning Editor at Bloomsbury Publishing and is responsible for the British Wildlife Collection. She recently took some time to chat with us about this excellent series; how it began, what makes it stand out from others of its kind and what we can expect from the collection over the next year. Keep reading for the full conversation with Katy, and browse the full British Wildlife Collection at nhbs.com.

Firstly, could you tell us a little bit about yourself; what you do and what your involvement is with the British Wildlife Collection?

I’m a Senior Commissioning Editor in the wildlife team at Bloomsbury Publishing. The British Wildlife Collection is one of the lists I’m lucky enough to look after; I’m responsible for signing up new books with authors and then seeing them through to the point at which they go off to the printer. Being immersed in these books means that I’m always learning something new: from the surprising fact that saltmarsh occurs as far inland as Staffordshire, through to how to successfully use a pooter to collect ants (without inhaling live insects!).

Published in 2012, the first title in the series was Mushrooms by Peter Marren. Could you tell us more about how the British Wildlife Collection initially came about?

Yes, the series was conceived by Andrew Branson who founded and, for 25 years, ran the publication of the excellent British Wildlife magazine. I believe that Andrew’s intention was to produce a series of seminal books that would capture the essence of the magazine in terms of being informative, well-written and thought-provoking whilst providing the author the space to explore their chosen subject in detail and develop the overall narrative.

For anyone unfamiliar with the books from this collection, how would you describe them and who do you consider to be their intended audience?

The series covers all aspects of British natural history and we encourage our authors to cover elements of ecology, history, management, conservation and culture; in other words, to explore the human relationship with their particular subject. The books are written in a way that we hope appeals to a wide readership, from academics and conservation professionals through to enthusiastic amateur naturalists and people who simply want to know more about the British countryside and its inhabitants.

All the books in the series are incredibly well-researched and comprehensive. How long does it typically take to go from conception to publication for a single title?

It varies depending on the author and their other commitments, but I’d say it typically takes around three years from the date the contract is signed until the book hits the shelves. I think our record for the shortest time is 13 months (I won’t tell you the longest!).

The books are beautifully produced and printed and, without exception, are authored by the most well-respected experts in their field. It is no surprise that they have rapidly become collectors’ items for so many British naturalists. What do you think makes these monographs stand out from others of their type?

The series is relatively new and is still evolving, but our ultimate aim is to build an indispensable reference source on all aspects of our wildlife by commissioning some of our finest writers and leading experts to write accessible and engaging books that are then carefully designed to incorporate hundreds of carefully chosen photographs and illustrations to bring the text to life. We hope that our readers come away from them, as one reviewer put it, ‘better-informed than ever, but also hugely entertained’. Oh, and they each feature a beautiful specially commissioned cover artwork by the peerless Carry Akroyd.

Finally, are you able to give us any information as to what subjects we can expect to be covered in the next year or so, or is this a closely guarded secret?

Hedges by Robert Wolton is coming out in February this year, and beyond that we’re excited to be welcoming Peter Marren back to the series with a book entitled Rare Plants. He’ll be followed by Trevor Dines with Urban Plants and David Goode with Bogs.

All books in the British Wildlife Collection are available from nhbs.com. Please contact our customer services team if you would like to set up a standing order for this series – this will ensure that you automatically receive each new title as soon as it is published.