Interview with Jan Collins: Bat Surveys for Professional Ecologists 4th Edition

Purple cover for Bat Surveys for Professional Ecologists with lots of images of bats.The 4th edition of Bat Surveys for Professional Ecologists: Good Practice Guidelines is the latest update of the Bat Conservation Trust (BCT) Guidelines and features new content on biosecurity, night-vision aids, tree surveys and auto-identification for bat sound analysis. Several key chapters have been expanded, and new tools, techniques and recommendations included. It is a key resource for professional ecologists carrying out surveys for development and planning.

Portrait image of the author Jan Collins

Jan Collins is the Head of Biodiversity at the Bat Conservation Trust and a former ecological consultant. Her fascination in bats began when participating in a bat biodiversity survey on a Vietnamese expedition in 1999 and she has worked in bat conservation ever since. She has been in her current role for the last 10 years and has played a central role in the editing and refining of the BCT Guidelines.

In this Q&A we had the opportunity to speak with Jan about some of the key aspects of the 4th edition of the BCT Guidelines and its consequences for ecologists.


What led you towards a career specialising in bats?

From a young age I was interested in being outdoors and engaging with nature. My studies followed this route, with a degree in Environmental Sciences and a Masters in Ecology and Management of the Natural Environment. I was first introduced to bats on a Frontier expedition to Vietnam in 1999 where we were trapping bats as part of a biodiversity survey. I was immediately hooked (what an amazing group of animals, with unique behaviours, and so very diverse!) and sought both voluntary and paid work involving bat conservation upon my return. I was an ecological consultant specialising in bats for over 10 years before joining Bat Conservation Trust in my dream job as Head of Biodiversity almost exactly 10 years ago now.

Grey long eared bat flying with wings spread at night with leaves in the background
Grey Long-eared Bat (Plecotus austriacus).

Could you tell us briefly about the work that the BCT does?

BCT is a dynamic, influential and growing national charity. We are the leading non-governmental organisation in the UK devoted solely to the conservation of bats and their environment. Our work represents the gold standard in bat conservation providing a lead for the rest of the world. We work to ensure that bat conservation is acknowledged as an integral part of sustainable development. Our work ranges from best practice guidance, advice and training through to engaging wider audiences so that we can get more people to understand the importance of bats and their conservation. A lot of our work also involves working with partners on the ground, local bat groups play a huge role in all aspects of our work. More information about all of our work can be found in the most recent annual review, found here or take a look at the Bat Conservation Trust Website here.

The 4th edition of Bat Surveys for Professional Ecologists: Good Practice Guidelines is due to be published in September 2023. What have been some of the key challenges in getting this ready for publication?

One of the key challenges for this edition was gaining consensus from the Technical Review Board on some of the recommendations. This included the continued use of transects for bat activity surveys and how tree surveys for bat roosts should be carried out, acknowledging the various limitations, and how approaches should be adapted to different types of projects. Another challenge has been how much to include on night vision aids such as infrared cameras. BCT are planning a kick-off meeting for a working group to develop separate night vision aid guidelines in the autumn (to include infrared and thermal imaging cameras) and therefore many aspects are still to be discussed and decided. We will also run a public consultation on night vision aids to help us to understand current practice and expectations from new guidelines on this topic.

Infrared camera setup in foreground pointed at a house in the background for bat survey
Infrared camera setup. Photo and setup by Richard Crompton.

How has the outlook for bat populations in the UK changed since the last edition of the Guidelines?

Data from the National Bat Monitoring Programme indicate that populations of the bat species we monitor in the UK are stable or recovering. However, it should be remembered that these trends reflect relatively recent changes in bat populations (since 1999 for most species). It is generally considered that prior to this there were significant historical declines in bat populations dating back to at least the start of the 20th century. This suggests that current legislation and conservation action to protect and conserve bats is being successful, and it is vitally important that this continues. Detailed information on trends for the 11 species monitored can be found in our National Bat Monitoring Programme reports here. We are also seeing signs of regional variations that deviate from the overall positive trend at the national level and we want to gain a better understanding of those. More data would help this process so we would encourage everyone to join in with one or more of the National Bat Monitoring Programme surveys. There are surveys that are suitable for anyone regardless of experience or equipment.

What are some of the key changes in the Guidelines?

A number of chapters have been expanded and new tools, techniques and recommended best practice incorporated. Below are a few of the changes but we will be developing a webinar in the coming months, detailing all of the changes so readers should watch out for that!

  • A new section on Biosecurity has been added to Considerations for Bat Surveys chapter, recognising that precautionary approaches are needed to protect both ecologists and bats from potential health risks.
  • Chapter 6 on Surveying Trees and Woodland for Bat Roosts has grown from five pages in the 3rd edition and now includes details of newer technologies and sources of information such as night vision aids, motion activated camera monitoring, The Bat Tree Habitat Key and the Bat Roost Tree Tag Project as well as updated guidelines for categorising the suitability of potential roost features.
  • Chapter 7 focuses solely on dusk emergence surveys, with dawn re-entry surveys removed as a standard approach due to the improved quality of emergence surveys with night vision aids and the variability in the time that bats return to their roosts.
  • Chapter 10 on Data analysis and Interpretation incorporates information on auto-identification systems and has a new section on data science which includes details on elements such as tidy data, minimal data requirement and data standardisation. There are also case studies to help illustrate key points.

How will the new Guidelines improve the way ecologists approach bat surveying?

The guidelines are the key resource for professional ecologists carrying out surveys for development and planning. The Biosecurity section in the 4th edition will ensure that ecologists carry out their work in a safe way, minimising health risks to both themselves and to bats. The latest edition acknowledges the constraints involved in surveying trees for bat roosts and offers different approaches to surveys depending on the nature, scale and timeline of the project. A new categorisation system for Potential Roost Features (PRFs) and trees reduces the subjectivity in initial assessments of trees. Dawn re-entry surveys are no longer recommended as there are questions around their efficacy for presence/absence and because emergence surveys can be vastly improved by the use of night vision aids. A stepwise approach is offered to those using auto-identification systems to analyse acoustic datasets and the section on Data Science aims to standardise approaches to data management. The guidelines also make reference to a wealth of resources that can be used to improve surveys and, in particular, interpretation of survey data.

Horseshoe bat hanging from a rock by its feet with its face facing the camera.
Horseshoe bat (Rhinolophus ferrumequinum).

Nocturnal survey equipment such as night vision, thermal imaging and infrared cameras are becoming more important in bat survey with the 2022 Interim Guidance Note providing clarification of their role. How will the 4th edition of the Guidelines impact the way ecologists conduct bat surveys using these types of equipment?

The 4th edition supercedes (but is consistent with) the Interim Guidance Note published in May 2022. It emphasises that surveys should usually be carried out with night vision aids and that if they are not used this should be justified in reporting, with reasons provided (e.g. at known roosts when bats are known to emerge early or in situations/locations with higher levels of natural or artificial light). Dawn re-entry surveys are no longer recommended as a standard approach because the use of night vision aids vastly improves the quality of emergence surveys, when used properly. The new guidelines state that a still shot must be taken at the darkest point of the survey to show the field of view and that appropriate illumination has been used. They suggest that the use of night vision aids can be used to reduce the number of surveyors but only if the cameras/lighting reliably match or exceed what a surveyor can achieve, with evidence provided. See the 4th edition for more information on use of this equipment!

BCT are planning a kick-off meeting for a working group to develop separate night vision aid guidelines (with much more detail) in the autumn (to include infrared and thermal imaging cameras) and therefore many aspects are still to be discussed and decided. We will also run a public consultation on night vision aids, to help us to understand current practice and expectations from new guidelines on this topic.


Bat Surveys for Professional Ecologists: Good Practice Guidelines is now available to pre-order from nhbs.com.

Author interview with Fiona Mathews and Tim Kendall: Black Ops & Beaver Bombing

Black Ops & Beaver Bombing: Adventures with Britain’s Wild Mammals is a captivating and entertaining deep dive into many of the mammals of Britain. Fiona Mathews and Tim Kendall explore mines inhabited by great horseshoe bats, go on overnight stakeouts in search of pine martens and travel from Scotland to the Isles of Scilly in search of their elusive subjects. This book puts animals at the heart of the story, revelling in their peculiarities and exploring the threats to their survival and the struggles that plague their conservation in Britain. The depth of knowledge, witty commentary and obvious enthusiam creates a beautifully written book that is difficult to put down.

Each chapter focuses on a different UK mammal, from wild boars and beavers to red squirrels and grey seals. Fiona and Tim explore the history, ecology and current conservation of these species, focusing on what is threatening them and what should or is being done to protect them. They do not shy away from giving criticism and sharing their frustration when the attitudes of policy makers stand against environmental protection and restoration. In search of answers to the problems that beset our wildlife, the authors reveal the wonder of creatures that are worth fighting for.

Fiona Mathews is a professor of environmental biology at the University of Sussex and the founding chair of Mammal Conservation Europe. Tim Kendall is a professor of English Literature at the University of Exeter and edited Britain’s Mammals 2018. They have kindly agreed to an extended interview with us, where we discuss the role of public education in conservation projects, how climate change may be impacting population recruitment in bats and seals, whether there is hope for the reintroduction of top mammalian predators, and much much more.


Could you tell us how you both became interested in mammals and what inspired you to create this deep dive into the mammals of Britain?

Tim: I grew up right next door to Devonport Dockyard in Plymouth. I was obsessed with spotting mammals as a child, but the only ones I ever saw were grey squirrels. When I was 8 or 9, I wrote a poem for school about seeing a badger, describing the joy and excitement I felt. It was all made up: I didn’t see a live badger until after I left for university. Luckily, I had the good sense to marry a mammal biologist and carry out fieldwork with her, which means that I’ve now seen all but three of our native mammal species.

Fiona: I’ve been interested in mammals for as long as I can remember, but I decided it should become my career after I went on my first bat walk. Black Ops and Beaver Bombing was partly inspired by my work reviewing the population status of Britain’s mammals and drawing up the Red List with the Mammal Society.

The preface, I-Spy, makes an interesting point about the interconnectedness of species as well as the focus of conservation: “If you want to save the barn owl, start by saving the field vole”. Do you believe the focus on ‘charismatic’ species, which are often higher up in the food chain, more often aids the conservation of prey species (such as by increasing or improving habitats) or harms these species by increasing the population of their predators? Should more conservation projects take a more ‘grassroots’ focus, by helping species lower down on the food chain?

We ignore the small species at our peril, and the fact that they’re almost all in decline is deeply worrying. Of course, wolves and wild cats grab the headlines, but if we don’t pay attention to the bioabundant species, everything across the food web is affected. We talk a lot in the book about the Scilly shrew, and if we’d thought anyone would read it, we’d have included a chapter on field voles.

In Chapter 3 ‘On the Trail of the Lonesome Pine Marten’, you discuss the extensive work done to win over the local people before the ‘recovery’ project began. Do you believe the public will become more and more open to these types of projects or will public education always need to play such an extensive role?

The general public is already becoming more welcoming, but the people who control landscapes are a small subset of the population, and they often have their own priorities that they believe (rightly or wrongly) will be damaged by reintroduction and recovery projects. One of the unfortunate things about rewilding is that it’s becoming as divisive as Brexit. We should all be working together to reverse the biodiversity crisis.

Wild mammals only make up between 2–4% of the world’s mammal biomass, decreasing by more than 85% since the rise of humans. In contrast, livestock makes up more than 63% of mammal biomass. How has this drop in wild mammal biomass impacted ecosystems and do you think we’ll be able to repair this?

High intensity livestock production is undoubtedly a massive contributor to biodiversity loss, but we mustn’t perceive farmers as the enemy. We need good farmers producing good food and getting rewarded for sustainable and wildlife-friendly production. Progress towards reforming agricultural subsidies has moved at glacial speed. Our government, like most across the globe, is much too heavily influenced by the lobbying of large agribusinesses which want to preserve the status quo.

In Chapter 5 ‘Hanging Out with Greater Horseshoe Bats’, you mention that Fiona is working on a long-term plan to re-establish the British population of the greater mouse-eared bat. Could you tell us a little bit more about how this process would work?

Fiona: Historically, the greater mouse-eared bat has been found in Dorset and Sussex, but with climate change we would expect its range to shift northwards. There are large and thriving colonies in Brittany and Normandy at the moment. We have to work out whether they will arrive in England under their own steam, or whether we need to give them a helping hand. Translocating bats isn’t easy. I’ve set up a working group at Eurobats to bring together experts looking at the best way forward.

You mention that the greater horseshoe bat population has stabilised and is now increasing in Britain. Why do you think this is happening?

The species is struggling across mainland Europe but it has responded well to a series of mild winters in England and Wales. Organisations like the Vincent Wildlife Trust have done a fantastic job to protect and improve roosts. We should remember, though, that greater horseshoes once had a range that extended to the east of England, and it’s estimated that there was a population of 300,000 at the end of the nineteenth century. The current population is about 4% of that total, which is a remarkable recovery from their lowest levels, but we need to be aware of shifting baseline syndrome before we celebrate the good news too enthusiastically.

Chapter 5 also mentions a new system for monitoring bats, which monitors the flight paths using radio receivers. Do you think that, by highlighting important flight paths, this system will be able to be used to reduce the impacts of new developments?

Although legislative protection means that we now know a lot about bat roosts in buildings, there’s not much point protecting a roost if the animals lack places to forage or mate. New techniques such as static radio tracking allow us to identify important sites and protect them for the future.

As cool weather in spring can have an impact on the growth and development of young bats, has climate change impacted juvenile survival rates and population recruitment?

We know that baby bats grow less well, with short- and long-term consequences, when weather conditions are poor in the breeding season. We would expect to see negative effects from climate change, and this is something we are currently looking at in a European-wide project. The challenge is that few researchers have datasets that span at least 20 years, which is the timeframe you need for analysing these long-term trends. So we’re also conducting other research on shorter timescales to assess whether mother bats are moving around their roosts to select different temperatures.

Chapter 6 ‘Tiggywinkle Goes Rogue’, mentions how ‘tidiness’ harms biodiversity. Do you think the growth in recent campaigns such as No Mow May will help to reduce society’s obsession with garden ‘neatness’?

Yes! It also requires us to hassle our local councils until we outnumber those strange people who complain about overgrown verges. Councils will take the path of least resistance, which should be to do as little ‘tidying’ as possible.

A common threat for several of the species you cover in Black Ops & Beaver Bombing is light pollution. Is there momentum at the moment that would see light pollution effectively combated in the coming years?

No, and the problem is worsened by the fact that lighting is getting cheaper and more intense. It’s frustrating because light is one of the few pollutants that simply disappears as soon as you flick a switch.

In chapter 7, ‘Who Cares What Colour the Squirrels Are?’, you mention that certain conservation strategies, such as the culling of grey squirrels to protect red squirrels, are unlikely to receive funding due to fears of adverse publicity. How much do you think conservation strategies are affected by how palatable they are to the public? Do you think this impacts the effectiveness of some conservation projects?

The biggest issue is the insatiable appetite for novelty over longevity. Nobody wants to fund routine maintenance. You may get grant money to reintroduce water voles, but will you still be funded to cull American mink a decade later?

As storms often lead to the deaths of many seal pups, will the increased occurrence of extreme weather due to climate change have a significant effect on population recruitment for seals on our shores?

We don’t know. In our seal chapter, we call the recovery of grey seals the great mammal conservation story of the twentieth century. Their numbers increased from about 500 during the First World War to well over 100,000, which makes up around 40% of the global population for the species. Storms and tidal surges can wipe out entire colonies of pups in any given season, but thankfully so far that doesn’t seem to have dented their population growth. Shout out to the amazing volunteers who rescue and raise pups for 6 months before they’re strong enough to be released!

The government recently released England’s first Marine and Coastal Wildlife Code, do you think this guidance will significantly help to protect wildlife such as grey and harbour seals from the impacts of visitors? 

Yes. It was long overdue, and it’s vital. Some people will remember seeing the footage of a runner deliberately chasing a colony of seals into the sea in Yorkshire. If that happens to a moulting seal or a pup, they can die. We’re all tempted to take selfies next to wildlife, but it’s rarely a good idea.

This book ends by asking why Britain cannot seem to accept mammalian predators in the way that European countries have. Do you think there is hope for change in attitudes that could see the re-introduction of species such as lynx, wolves and bears within the coming decades?

We end the book in the Abruzzo National Park, 90 minutes drive from Rome, where there are now about 11 active wolf packs and 50 bears. The Cairngorms is ten times bigger, and has roughly the same human population. So much for the argument that we’re a crowded island! So we could and should reintroduce these apex predators, but we have to deal with misinformation from (for example) the current President of the National Farmers’ Union, who claims that lynx would pose a threat to ramblers. No wild lynx has ever attacked a human. Cattle, on the other hand, kill 8-10 people a year in Britain.

Do you have any future plans that you could tell us about?

We’ve just started a podcast, ‘Mammals R Us’ (https://rss.com/podcasts/mammalsrus/). We have very different working habits, so our big question is: can we finish another book without getting divorced?


Black Ops & Beaver Bombing by Fiona Mathews and Tim Kendall was published by Oneworld Publications in April 2023 and is available from nhbs.com.

Author interview with Michael Stephen Clark: The Fragmented World of the Mongoose Lemur

The Fragmented World of the Mongoose Lemur is a book about a critically-endangered species, fractured ecosystems and the global war on nature. It is a compelling story in which author Michael Stephen Clark reveals the fundamental paradox of the mongoose lemur’s natural history.

In this post we chat to the author of this important and timely book about his work with mongoose lemurs, the problems withthe way that conservation is reported in the mainstream media and his hopes for the future given the alarming loss of biodiversity and rate of extinctions we are currently experiencing.


Author Michael Stephen Clark

Your book is described as a modern natural history, but it also conveys a larger message that is of relevance to our relationship with all wild species. Why did you choose the mongoose lemur as your focal species – what is it about this primate that makes it suitable for illustrating more widespread issues?

I first became interested in the mongoose lemur when I worked with an aged and overweight pair at London Zoo in the 1980s and 1990s. They were being kept off-exhibit, which I thought was inappropriate given their endangered status. In due course, we were able to move them into a larger, outdoor enclosure, but only after an operation had been performed on both of them to remove excess adipose tissue. It was all very invasive, but the animals were much fitter, healthier and happier as a result. Shortly afterwards, I discovered that the captive breeding programme for the species lacked co-ordination, so I established the first international studbook for the species in captivity. There wasn’t a great deal of information about them in the literature and relatively few researchers and institutions had sought to remedy that. It quickly became clear to me that the mongoose lemur was fast becoming undervalued and potentially overlooked as an endangered species. I’ve been concerned about them ever since, probably because my sympathies often lie with the ‘underdog’.

I’d long been aware that the mongoose lemur could illustrate very directly the profound disruption caused by human alteration of the natural continuum. Human attitudes to wildlife contain a raft of contradictions that result in contradictory outcomes. This is particularly true of the mongoose lemur. In common with virtually all lemurs, it is a critically endangered species that exists in compromised and/or fragmented habitat in the wild. Perversely, however, it appears (on the surface) to be more numerous and relatively safe as a naturalised alien species on the Comoros Islands of Anjouan and Moheli. Lemurs in captivity are generally viewed as a conservation priority, yet the mongoose lemur ranks lower in importance than some of the more enigmatic lemur species. Already, without digging too deeply, we can see that the mongoose lemur has experienced displacement at every point where it has come into contact with humanity.

Perhaps, more alarmingly, the mongoose lemur has unwittingly become a prima-facie example of something disquieting; namely, a growing acceptance that nature will be allowed to persist only in places of our choosing. It’s a situation that has led to questionable decisions around relocation, translocation, restoration, rewilding, and re-introduction. To my mind, these things invite compromise, not least where species such as ring-necked parakeets in England and mongoose lemurs on Anjouan are inter-changeably considered wild, free-living, native and/or naturalised. These compromises ominously signpost natural systems by human design. Presently, the momentum is behind stand-alone programmes, projects and initiatives that are, of necessity, limited in scale. This may well lead to the ultimate breakup of biodiversity and an end-point where our natural world has become little more than a ‘zoo in the wild’.

Do you think that the way conservation and the plight of endangered species are reported by the mainstream media is problematic? I’m thinking, for example, in terms of how the public, conservationists and governing bodies might go on to perceive the state of the natural world as a result?

I actually think it’s a massive problem for conservation. Once upon a time, the adage was ‘there’s no such thing as bad publicity.’ In recent years, the mainstream media has shifted away from the substantial to the superficial reporting of wildlife conservation matters. Social media and the wider online milieu all feed into that, of course, but far too many stories are either half-told, poorly-told, or just plain misleading. It’s actually worse than not telling the story at all. This is especially disruptive for governing bodies and NGOs that can so easily find themselves painted into a corner by media-generated hubris. I’m not sure that a proliferation of conservation news portals, websites, blogs, podcasts and influencers is useful either. It is more likely that conservation stories will become even more distorted, diluted and diffuse as result.

I think the effect of all this on the public, among conservationists, and within governing bodies is potentially quite dangerous. It could make identifying priorities and taking effective action a minefield of conflicting imperatives. The recurring controversy over the Ramsar site at Coul Links in Scotland is a good example of media hubris threatening not only to drown out voices of reason, but also to trample the letter of the law into the ground. It took the combined firepower of a self-generated media campaign by a coalition of Wildlife Trusts and NGOs to beat back an existential threat to unique (and legally protected) dune habitat. I suspect that such coalitions will have to establish permanent media entities if they are to fight the war on nature across several (media) fronts.

Given the alarming rate of extinctions and global diversity decline, what would you like to see happen, both on a local and an international scale, to begin dealing with this? And would you describe yourself as broadly hopeful or pessimistic about this crisis?

I’m afraid that I am neither optimistic nor pessimistic about the future. I know it’s a cliché, but I think the future is already here. The important decisions now (as ever) will lie with young people, many of whom are not only well-educated and eminently qualified, but savvy as well. I recently rejoined Linkedin after a long absence and the sheer number of extremely active, professional wildlife conservationists around the world is really quite dazzling. In the short-term, issues around job security, personal security, career development, and establishing substantial permanent facilities in the field need to be addressed.

Frankly, I think that’s primarily the top-down responsibility of pan-national institutions such UNEP and the World Bank. The scale of accelerated extinction events forewarns us of a looming biodiversity crisis that could very easily become an existential threat in just a couple of generations. It requires a global response and the money really has to start flowing from global institutions if we are to empower and support our highly motivated international conservation professionals. To my mind, food security, health, climate, biodiversity and species conservation are equal priorities. I don’t understand how this became a hierarchy of aspirational goals because the response they all require is equally urgent.

In the medium term, it is not news to say that we have to invest in future generations, but ‘hope and inspiration’ aren’t enough. I think that young children generally receive a good grounding in natural history in their pre-school and primary school years. With a little support, that can happen in a developing country as easily as it can in the UK. But, young people need to have pathways they can follow in order to learn more. From there, they can gain a rounded understanding of natural history, which would inevitably include an elementary form of conservation biology. I’m a bit surprised that some of the more prominent NGOs aren’t lobbying for this with a draft curriculum. Nature conservation education for school-age children has to graduate from photocopied fact-sheets and elevate itself as the most valuable ‘eco-system service’ available.

As a zoological professional you have studied and worked in several esteemed organisations including the University of Bristol, London Zoo and Oxford-Brookes University. What do you consider to be the most significant highlights of your career so far?

Anyone who has ‘hands-on’ experience of working with wild animals in the field and/or captivity will tell you it’s a life full of great highs and lows. It doesn’t do to dwell on the lows so I’ll concentrate on things that I considered achievements. As I explain in my book, the time I spent on Anjouan in the Comoros Islands was a very special career highlight. It was a great privilege to be trusted with the capture and care of Pteropus livingstonii, one of the rarest bat species in the world and, of course, to encounter free-living mongoose lemurs first-hand. I’m especially gratified that several papers, features and articles that I authored or co-authored were published in the course of my work at the Zoological Society of London. These included my account of mongoose lemurs on Anjouan, the jointly written history of black-footed penguins at London Zoo, and a report on breeding Leadbeater’s Possum at London Zoo. Getting up close and personal with urban foxes and badgers in the course of fieldwork for Professor Stephen Harris at Bristol University was memorable, and I very much enjoyed my brief role as a visiting lecturer at Oxford-Brookes University. It was quite an honour to be there at the beginning of the now prestigious Primate Conservation MSc course.

The Fragmented World of the Mongoose Lemur is, in large part, a species-specific natural history – a format that is perhaps less popular than it has been in previous years and decades. The book was also published independently. How did you find the process of planning, writing and publishing the book and what were the main challenges you faced?

It’s correct that the species-specific natural history books have fallen out of favour with mainstream publishers, but I’m not convinced that readers feel the same way. In (apparently) normal circumstances, the life history of a species will appear to be relatively constant. In natural history classics such as David Lack’s ‘Life of the Robin’ and Sarah Churchfield’s ‘Natural History of Shrews’, the subjects live in a self-contained world that is full of challenges, yet seems immutable. The world today, in the age of the Anthropocene, rarely allows for ‘normal circumstances’. Wild animals will live, prosper, suffer, die or become extinct largely at our discretion, which means that their life histories now have to take account of the rapid, ongoing changes wrought by hyper-accelerated, unregulated human activity. That’s why I think it’s valid to revisit species-specific natural history writing and place it in a much wider context.

There was a time when my book would have been titled ‘The Mongoose Lemur in its World’, but that world no longer exists. It more accurately lives in a fragmented world of our making. This is happening to species the world over and those that are able to adjust may live, while those that can’t will surely disappear. We need to understand what makes the difference between the two potential outcomes in order to construct a meaningful response. I think this can emerge from species-specific natural histories that draw from a deeper well of scientific, philosophical and practical resources. I think it’s too often wrongly (and condescendingly) assumed that readers either can’t or won’t respond to stories that cast the net widely and speculate, as I have done, about ‘Propects and Perspectives’.

I’m fairly comfortable now with the process of independent publishing and I’ve never seen it as a poor relation to using an established publishing house. I would describe it as a slow, gentle learning curve rather than a steep and arduous one. The early stages of planning and writing are the most fun, but luckily I quite like editing, proofing and polishing too. I’m quite experienced in that regard but I find that editing your own work means that you really can’t rush it. Familiarity breeds oversight. I usually keep sharing with others to a minimum, but for this book I got a lot of help on specific chapters. I needed it too, and their input was invaluable as the acknowledgements section of the book will attest.

If I had to identify a single challenge that was really testing, I would say that maintaining a consistent, balanced style of writing throughout gave me the most sleepless nights. I wanted to hold the reader’s interest with an engaging narrative but still maintain a firm grip of the factual material. I also wanted the book to be a bit provocative. I always think I’ve read a good book if it leaves me with as many questions as it does answers.

Finally, what is in store for you next? Do you have plans for further publications?

At the moment, I’m doing the preparatory work for events in support of the book. I’ll be using my original colour slide transparencies from Anjouan to illustrate my presentations and I recently bought a very old colour slide projector for that purpose. It’s a bit quirky, I know, but I think it will make things a bit more interesting.

In terms of writing and publishing, I have a couple of things of my own in development, but I’d be interested in collaborating with others to produce further modern natural histories. I’d like to keep the focus on primates, mainly because they’re exposed to lots of different pressures in a variety of situations where they come into close contact with us humans. Their stories challenge us to think about the relationships we have with nature generally and how we are going to rescue biodiversity from our worst excesses. Species such as pygmy slow loris, Cat Ba langurs and Kirk’s red colobus spring to mind, although there are countless others.

Print-on-demand and increasingly accessible typesetting software mean that there are few obstacles to producing books like this. If there is a problem then it is persuading authors to enter a publishing relationship that is unconventional, yet fairer and more equable than the conventional model. Publishing independently also means that your book need never go out of print, something that frustrates many published writers who find themselves out of contract.


The Fragmented World of the Mongoose Lemur by Michael Stephen Clark is available now from NHBS.

In The Field: Batbox Baton Bat Detector

The Batbox Baton is an economical and user-friendly bat detector ideal for newcomers to bat detecting and bat detecting enthusiasts alike. The Baton is perhaps one of the most simple and easy-to-use bat detectors on the market, so simple that it can be operated with a single button. With simplicity often comes sacrifice, but not in the case of the Baton. This device uses technology called frequency division which enables the user to monitor all ultrasonic frequencies between 20kHz and 120kHz at once by dividing the frequency by a factor of 10. If a bat calls at 50kHz, for example, a 5kHz form will be played through the speakers. This means no tuning is required and the user is not at risk of missing any bats by being tuned to the wrong frequency.

We took out a Batbox Baton to a rural lake in Hampshire at dusk on a dry August evening. The detector comes preloaded with a battery, and with a flick of the single On/Off button we were listening to bat calls in a matter of seconds. The detector is extremely lightweight, ergonomic and compact, making it easy to carry into the field. The calls of (what we believe were) Soprano Pipistrelles were divided down to an audible frequency and we could hear multiple individuals calling and hunting above us. It is worth noting that species identification can be more difficult without a frequency display screen, especially if the user has less experience in hearing calls in frequency division or if they are unable to compare with other bat calls. We found the Baton a very useful tool for listening to bats for pleasure and the lack of a screen or tuning dials means you can focus your eyes above and watch the bats as they fly and hunt. 

Should the user wish to get a bit more out of their bat detecting experience, however, the Baton does provide options. The Baton has a ‘Line Out’ socket, and when connected to a laptop with a soundcard via a stereo lead, and used in conjunction with the free BatScan analysis software compatible with Windows only, real-time sonograms can be viewed in the field allowing detailed analysis and species identification. 

The Baton’s Line Out socket can also be used with a digital audio recorder, such as a H1n Handy Recorder, and calls can be recorded for future analysis using the same BatScan software. It should be noted that if the user wishes to listen to calls through headphones, this cannot be done through the detector itself but only via the audio recorder. The use of a recorder and further analysis with BatScan software allows the user to gain a detailed understanding of call structure and species identification, and further their enjoyment of bat detecting. 

Whether you have been enjoying bat detecting for years, or you are just looking to start out, the Batbox Baton will have something for you. It is an economic and versatile option that we would not hesitate to recommend.


The Batbox Baton Bat Detector can be found here. Our full range of bat detectors can be found here.

If you have any questions about our range or would like some advice on the right product for you then please contact us via email at customer.services@nhbs.com or phone on 01803 865913.

Owl Pellet Dissection

Owl pellets contain all of the indigestible parts of the prey. Image by Gail Hampshire via Flickr.
What is an owl pellet?

Owls feed on a variety of prey; most commonly small mammals but also birds, frogs and other small animals. These prey items are consumed in their entirety and, while the flesh is digested by enzymes, the owl is unable to digest the harder parts of the body, including the teeth, bones, fur or feathers. These indigestible parts are regurgitated as a pellet. Unless they are very fresh, pellets are dry, light and odourless.

In this article we will look at where to find owl pellets and how to tell which species of owl they came from. We will also provide some tips on how to dissect a pellet, how to group the bones into types, and how to identify some of the main species of small mammal that you will find in pellets in the UK.

Where to find owl pellets

Owl pellets can frequently be found wherever owls nest or roost. Good places to search are at the base of tall trees within woodland areas, or in barns or outbuildings where owls are known to roost. Please be aware that you must not disturb breeding or roosting owls in order to collect pellets. Barn Owls in particular are protected in the UK by law under the Wildlife & Countryside Act 1981 and their breeding sites must not be disturbed under any circumstances.

If you don’t have any luck finding your own pellets or don’t have access to places where you might find them, there are several places online where you can order some. Reputable sources in the UK include the Barn Owl Trust and the Suffolk Owl Sanctuary.

What species of owl is my pellet from?

There are five resident species of owl in the UK: Barn Owl, Tawny Owl, Little Owl, Short-Eared Owl and Long-Eared Owl. All of these species produce pellets that are relatively easy to tell apart, particularly if you also know the habitat where they were found. Below is a brief guide to their main characteristics.

Barn Owl: pellets usually measure 3-7cm in length and are rounded at both ends. They are fairly dark in colour and have a smooth surface.
Tawny Owl: pellets measure 2-5cm in length and are narrow and bumpy, often having tapered ends. Greyish in colour and sometimes furry looking.
Little Owl: pellets are fairly small measuring only 1.5-2cm in length. Long and narrow with a soft crumbly texture.
Short-Eared Owl: pellets are fairly large, measuring 3-6cm in length. Narrow with one rounded end and one tapered end. They are grey and smooth and very lightweight.
Long-Eared Owl: pellets measure around 2-4cm and are narrow and bumpy. Usually grey in colour.

The most common pellets you will find in the UK are from Barn Owls.

Barn Owl pellet with mounted needle and fine pointed forceps.
How to dissect an owl pellet

There isn’t much equipment you need to dissect an owl pellet, but a few items will make the job a bit easier:

Mounted needle: this is useful for teasing out fur from around the bones, and moving around delicate specimens. A cocktail stick or needle pushed into a cork will also do the trick.
Forceps/tweezers: helpful for picking up bones and particularly for removing fur from inside skulls. Forceps with a fine point are best.
Magnifying glass/hand lens: a small magnifier will allow you to get a closer look at the bones that you find. Jaw bones in particular are very useful for identifying the species and a magnifier will help you get a better look at the arrangement and structure of the teeth.
White paper/card and glue: it can be helpful to arrange your bones by type onto a sheet of white paper which you can then write on when you have decided what they are and who they belong to. If you would like to make a permanent ID aid you can also glue them onto a piece of card and add permanent labels.

How to identify the contents of an owl pellet

The first thing you will need to do is to tease apart the pellet and separate the bones from the fur and feathers that are holding it all together. To begin with it is easiest to gently break the pellet into several smaller sections then work on each of these in turn. Use your fingers as well as the forceps to carefully tease apart each section, removing any bones and placing them to one side for identification. If your pellet is very hard and dry, try soaking it in water first to soften it.

Once you have all of the bones from your owl pellet, try to group them into types on your sheet of paper. The most common bones you will find are the following:

  • Skulls: for mammals, this consists of the top part of the skull and upper jaw, along with the lower jaw, although this is likely to become detached once you have cleaned all of the fur and other material from inside. For bird species this will include the upper and lower parts of the beak.
  • Back legs: includes the thigh bone (femur) and the lower leg bones (fibula and tibia)
  • Front legs (arms or wings): includes both upper (humerus) and lower (radius and ulna) bones
  • Hip bones
  • Shoulder blades (scapula)
  • Back bones (vertebrae)
  • Ribs

The image below illustrates typical examples of each type of bone. You can also download a useful bone identification sheet from the Suffolk Owl Sanctuary website.

 

The most useful part of the skeleton for identification is the skull and jaws or beak. Bird skulls will obviously be very distinct from those of mammals due to the presence of the beak, so these can immediately be separated out. For the remaining mammal skulls, however, we will need to take a closer look at their lower jaw bones and teeth.

In the UK the most common small mammals you will find in owl pellets are voles, mice and shrews. It is very easy to distinguish which of the lower jaws belong to shrews as they have a continuous line of teeth from the front to the back of the jaw. This is because shrews are insectivores and chew their food, much the same as we do. Voles and mice, however, both gnaw their food, and have a big gap between the long front tooth and the back teeth.

To tell the difference between voles and shrews, we need to take a closer look at their back teeth. Voles have teeth with distinctive grooves down the sides.  In those of a field vole, the grooves run all the way down the side of the tooth. There is also no obvious root. Bank voles have grooves which only run part-way down the side of the tooth and they have two obvious roots, similar to those of a human tooth. The back tooth from a mouse jaw is much smaller when compared to a vole and its structure is much more similar to that of a human tooth. It also has two roots. This sheet from the Barn Owl Trust has a great illustration of the various small mammal lower jaws with size guidelines to help with identification.

Hopefully this article has been a useful introduction to owl pellet dissection and the identification of some of the most common prey species contained within them. If you want more help with identifying all of the bones in your pellet down to species level, the guides listed below are invaluable. Once you have categorised all of the bones you can attach them to a piece of card with permanent labels or arrange them to create a complete skeleton of each species.

Finally, don’t forget to wash your hands well when you have finished your dissection. Any pellet remains can be safely composted.

Further reading

Guide to British Owls and Owl Pellets

This fold-out chart includes colour paintings of the five species of owl permanently resident in the British Isles, shown both perched and at rest. Also included are illustrations and written descriptions of the different pellets that may be found, and a systematic identification key to their contents, including complete skulls, jaws, teeth and other recognisable bones and animal parts.

 

The Analysis of Owl Pellets

This booklet will not only enable you to identify what you find in the pellets of British owls, but also shows how the data may be usefully presented and how to estimate the actual weight of food the birds have eaten.

 

In The Field: Elekon Batlogger S2

The Batlogger S2 is a compact passive recorder manufactured by Elekon. This all-in-one static bat detector and ultrasonic recorder is designed to be left unattended in the field over several nights to survey and monitor bats. The S2 is operated solely via Bluetooth and the BATLOGGER Control App (available on iOS and Android). It is small, weighing only 138g and measuring 132 x 72 x 35mm, but despite its size, the S2 is robust. It is waterproof and replacement microphones are also available, handy if the original microphone becomes damaged or loses sensitivity. 

Elekon has designed the S2 to be easy to use and lightweight, and built to withstand fieldwork conditions. We took the opportunity on a warm evening in mid-May to test the S2’s ability.

How we tested

The S2 was set up in a hedgerow in South Devon, close to a small known roosting site.

We connected the S2 to the BATLOGGER Control App on an iPhone via Bluetooth. Once connected, the S2 determines the dusk and dawn times using the GPS location from the phone/tablet, and suggests these as automatic trigger times. We selected this automatic time window, but, you can choose and customise your own and set multiple time windows as needed. 

The S2 is full spectrum with a range of 10-150kHz and a sample rate of 312.5 kHz. The default sensitivity is balanced, and we adjusted the sensitivity to ‘high’ using the App. This may lead to several unwanted calls in busier environments; however, it also ensures that it is triggered by most types of bat call, including social calls, which can sometimes be missed. 

Once collected from the field after one night of deployment, the recordings were downloaded from the S2 to a computer using the USB-C to USB-C cable (if you do not have a USB-C port, you’ll need an adaptor). Helpfully, the S2 is charged using the same USB-C cable – a single charge provides 100 hours of power!

The recordings are stored on an internal microSD card, and the S2 generates two file types: an audio file (.wav) and a recording information file (.xml). The audio files allow you to listen to your recordings through bat call analysis software and the information files store important metadata such as date, location, recording time, and device settings.

What we found

The BATLOGGER Control App shows you the number of sessions recorded – the high sensitivity triggered 192 audio files over one night. We used the BatExplorer software to manage and view the S2 recordings. The software has key features such as automatic bat call detection, making sorting files very easy, and it also provides suggestions for species identification. 

Of the 192 audio files, 40 of these identified the common pipistrelle (Pipistrellus pipistrellus). The default S2 sensitivity is ‘balanced’, this may have led to a smaller number of unwanted files (which recorded sounds other than bats). But we did not want to miss a bat call and the BatExplorer software allows you to quickly filter the unwanted files.

The echolocation frequency for common pipistrelles is approximately 45kHz, and the below images show an example of the spectrogram and call measures from a common pipistrelle recording taken at dusk and the information popout that BatExplorer produces, detailing the automatic analysis that the software carries out. 

Below is an audio clip with its accompanying spectrogram of a common pipistrelle taken at dusk. The BatExplorer software allows you to customise the playback and how the spectrogram can be viewed. 

Our opinion

The physical design and key features of the S2 makes surveying bats a simple task. 

The S2 truly is discrete and lightweight, making it easy to set up in the field, and once deployed the battery life will allow up to 10 (10 hour) consecutive nights of surveying. 

The setup through the BATLOGGER Control App is straightforward. The S2 conveniently uses the GPS location on your phone/tablet to determine dusk and dawn which benefits the accuracy of the recording schedules. You can choose your settings at a click of a button, and the instruction manual is clear and accessible for any help needed.  

The only limitation we found is that you cannot access the recordings straight from the App. However, transferring the files across to your computer allows you to listen to and analyse the audio files with ease. The BatExplorer software (available on a 30-day free trial) enhances analysis as it allows you to review, manage, and organise your recordings. 

The S2 is an impressive bat detector, and it is an ideal choice for professionals and ecological surveyors.


The Elekon Batlogger S2 can be found here. Our full range of passive full spectrum bat detectors can be found here.

If you have any questions about our range or would like some advice on the right product for you then please contact us via email at customer.services@nhbs.com or phone on 01803 865913.

 

The NHBS Guide to UK Deer Identification

Deer are among the UK’s most elegant and familiar mammals and sightings of them in their natural habitat are always special moments, however these encounters can often be fleeting, and our views obscured.

This ID guide covers all of the native and non-native deer species that are found in the UK, and describes the key features to look out for to aid in their identification.

Deer are hoofed ruminants that comprise the family Cervidae. They are naturally found across Europe, Asia and the Americas and can be divided into two subfamilies, differentiated mostly by their bone structures. Most familiar in the UK are the Cervinae or old world deer subfamily, which includes the red, sika, fallow, Chinese water and Reeves’ muntjac. The Capreolinae (new world) sub family includes the roe deer as well as elk, reindeer and all the species found across the Americas.  

Of the six species found in the UK, only the red and roe deer are truly native, although fallow deer were thought to have been first introduced to Britain in the 11th Century from the Mediterranean region, so are long established. Three other species, the sika deer, Chinese water deer and Reeves’ muntjac are all more recently naturalised within the UK.

Identification of deer can be straightforward in some situations, but some species are similar and, when not seen well, identification can be a challenge. Two of the best features to focus on for identification are the rump and the antlers, if they are visible. Except for reindeer (caribou), in which both sexes grow antlers, and the Chinese water deer and musk deer, which lack any antlers, all male (stags) deer usually grow antlers, which they use in battles to access females (hinds) during the rut. Antlers are unique to deer and a great tool to look at to identify different species. However, they are shed every year after the rut, so although a striking feature, the rump pattern of deer is perhaps the most reliable feature to use for identification.

Red Deer (Cervus elaphus)
Red deer by caroline legg via Flickr

Distribution: Widespread in Scotland and abundant in the Highlands and Islands. Elsewhere populations occur in Cumbria, Lancashire, the Peak District and Pennines, Exmoor and the Quantock Hills, the New Forest, and East Anglia. There are also small populations in Wales and Ireland.

Head and body length: 1.6–2.6 metres for a male and 1.7–2.1 metres for females

Height at shoulder: 1.14–1.22 metres.

What to look for: Both our largest species of deer and land mammal, the magnificent stags can weigh in at around 200kg making them an impressive and noticeably large species. Look for their reddish-brown coat that lacks any spots or delineation of colour. Only their rumps and tails feature a paler buff cream colour. Another characteristic of red deer are their elongated faces and large ears.

Their favoured habitat is woodland, although in Scotland they have adapted to live year-round in more open treeless landscapes. Grasses make up the bulk of their diet throughout the year, but they will also browse on a wide range of shrubs, young trees and bark, brambles, bracken and heathers. Sika deer are the most similar looking and the two species have hybridised in several regions. Sika have white spots in summer and darker brown coats in winter with shorter faces. Hybrid red and sika deer tend to resemble smaller darker red deer than they do sika.

Antlers: The antlers of mature stags are wide and branching with usually 8 sets of points per antler that curve upwards and sometimes in on themselves. Younger males have short unbranched, straighter pointed antlers.

Rump: A soft creamy colour with a very short russet-brown tail.

Sika Deer (Cervus nippon)
Sika dear by Chris Parker via Flickr

Distribution: Sika deer are native to Japan, Korea, Taiwan and far eastern Asia, but escaped into Britain from collections in 1860. Since their initial introduction in 1860, they have naturalised and spread to many regions. Population strongholds include Dorset and the New Forest, Lancashire and Cumbria Northern England, the Scottish borders and the Highlands.

Due to their genetic similarities to red deer, sika deer are thought to have hybridised with the native red deer in several regions, particularly in the Scottish Highlands. Sika prefer to keep to woodland cover more than red deer which have adapted to feeding in more open habitats. 

Head and body length: 1.2–1.9 metres for a male and 1.1–1.6 metres for female.

Height at shoulder: 1.07–1.22 metres.

What to look for: Sika are very similar in appearance to red deer and the two species do interbreed in many regions. They are noticeably smaller than red deer and in summer they have white spots on their coats, but thick and often dark (sometimes almost black) coats in winter.

Their diet and feeding habits are very similar to red deer, with grasses and heather making up the bulk of their diet, but they will also browse on both deciduous and coniferous trees, gorse, holly bark and acorns. They are however generally less social than red deer and outside of the breeding season, both males and females can be solitary with females forming only small herds with young.

Antlers: Similar to those of red deer, but thinner, lighter coloured and less complex with usually only 4 points per antler.

Rump: A conspicuous white rump patch with a dark edge and a short white tail with a single thin dark dorsal stripe along its length.

Roe deer (Capreolus capreolus)
Roe deer by caroline legg via Flickr

Distribution: A woodland specialist that’s rarely found far from some woodland cover, although they are increasingly using hedgerows and scrub as cover within more agricultural and urban landscapes. They are widely distributed throughout the UK but absent from Ireland, with the greatest population densities found in Scotland.

Head and body length: 0.95–1.25 metres

Height at shoulder: 0.6–0.75 metres

What to look for: A medium sized lightweight deer with a long neck and uniform brown coat. Other features that distinguish them from the larger deer include shorter muzzles and a clean white rump patch. In summer their coats turn a rich reddish brown and appear sleeker, while in winter, the coat turns a thicker and dark more peanut brown. 

They are mostly solitary but sometimes form small family groups with young, particularly during the winter. They browse a wide variety of trees, shrubs and herbs including bramble, heather, and rosebay willowherb but during the autumn will also feed on the ground on fruits, acorns and occasionally fungi.

Antlers: Short and mostly vertical, they are rarely much taller than the head, with only 2 or three points per antler. With a close view, the antlers can often appear particularly velvety or crusty at the base, depending on the season and growth stage.

Rump: The patch varies between the sexes, but both have a clean white patch and no visible tail. Males have a kidney shaped white patch, whereas females have more of a round shape.

Chinese water deer (Hydropotes inermis)
Chinese water deer by Nick Goodrum via Flickr

Distribution: The Chinese water deer is a native of eastern China and Korea but has formed a naturalised population in England after escaping from Woburn Park in Bedfordshire towards the end of the 19th Century.  They’re found throughout most of East Anglia and a more scattered population in the southeast-England where there is suitable habitat. The strongholds include the Norfolk broads and Cambridgeshire fens. The population trend is increasing, with over 1500 individuals and their distribution is also expanding. Interestingly, it is thought that due to population decline in their native range, the British population may now represent a significant part of their world population, despite not being a native species. 

Head and body length: 1 metre

Height at shoulder: 0.7–0.95 metres.

What to look for: A small and uniformly light brown (sometimes greyish) coloured deer with large, rounded ears and a distinctive black nose. They are strongly associated with freshwater marshland habitats where they feed on coarse grasses, reeds, herbs and aquatic vegetation. Chinese water deer are solitary and secretive, preferring to keep close to cover and use both woodland close to wetlands and reedbeds. They will occasionally forage in farmland but prefer to keep close to cover.

Chinese water deer by Nick Goodrum via Flickr

Males have impressive and prominent downward pointing tusks instead of canine teeth that can be seen with a close-up view. These tusks are used during the rut, mostly for display purposes between rival males and to impress females. They are quite a distinctive looking deer with a more unusual almost bear-like face, although their secretive nature means that obtaining good views can be difficult.

Antlers: This species lacks any antlers.

Rump: Their rear and short tail is the same pale brown colour as the rest of their coat. 

Reeves’ muntjac (Muntiacus reevesi)
Reeves’ muntjac by Peter Trimming via Flickr

Distribution: The Reeves’ muntjac is also native to China and again its UK population derives from escaped individuals from Woburn Park in Bedfordshire from 1901. They are now abundant and found throughout southern, eastern and central England spreading rapidly into southwest-England, Wales and southern Scotland. Since 2000, a population has become established in Ireland and Northern Ireland.

Head and body length: 0.9–1 metres

Height at shoulder: 0.45–0.52 metres

What to look for: A very small, robust and stocky deer that often appears to have a hunched-over appearance due to its short neck, arched back and tendency to walk with its head facing down to the ground. The coat varies from a deep russet brown in summer to a greyer and paler brown in the winter. The face is short and squat with a characteristic set of black stripes creating a V shape on the top of the head. Male muntjac also have slightly protruding tusks but they are much shorter than those of Chinese water deer and rarely visible without a close view.

Muntjac favour dense undergrowth within both deciduous and coniferous woodland but will also thrive within urban environments with suitable cover. They browse woodland leaves and flowers during the spring and summer including some scarce woodland ground flora species. During the autumn and winter their diet switches to nuts, fungi and grasses.

Antlers: A single very short point that curves back.

Rump: The rear patch and tail are the same reddish-brown colour as the back. The tail is short but when alarmed it will often raise the tail revealing the white underside.

Fallow Deer (Dama dama)
Fallow deer by Heather Smithers via Flickr
Female fallow deer by Steve Slater via Flickr

Distribution: Introduced to Britain for hunting in the 11th Century and to Ireland in the 13th Century from the Eastern Mediterranean, the species is now found throughout England, with more scattered populations in Wales and Scotland.

Head and body length: 1.45–1.55 metres for a male and 1.30–1.45 metres for a female.

Height at shoulder: 0.7–0.95 metres

What to look for: Fallow deer are quite variable in their appearance due to their wide range of pelage (hair). Typically, most individuals have some conspicuous white spots on pale brown, fawn or gingery coats in the summer and then dark brown coats with only faint or no spots in the winter. There is a great range of variation within this species though with some individuals and populations remaining very dark or very pale throughout the year, some of which can be melanistic. Fallow are medium sized deer that are very social, often forming large herds that remain together throughout the year.  Due to their numbers, they also leave conspicuous signs of their presence such as runs and tracks in frequented areas.

Their preferred habitat is open deciduous woodland but will also use farmland and woodland edge habits if there is cover close by. Fallow deer are grazers with grass forming most of their diet, although they will also eat nuts and browse on heather, holly and some deciduous trees to a lesser extent.

Antlers: Very large, broad and palmate shaped with numerous spikes.

Rump: A clean white rump with a dark outside edge. The tail is long and appears mostly black because of a long black stripe that runs down the course of its length, but the underside is clean white.

Further Reading:

Britain’s Mammals: A Field Guide to the Mammals of Great Britain and Ireland

£17.99

 

 

How to Find and Identify Mammals [Revised Edition]

£11.99

 

 

 

Mammals of the British Isles: Handbook, 4th edition jacket image

Mammals of the British Isles: Handbook

£34.99

 

 

 

A Guide to British Mammal Tracks and Signs

£3.75

 

 

 

Sika Deer

£4.99

 

 

 

Chinese Water Deer

£4.99

 

 

 

Scottish Red Deer and Their Conservation

£27.99

 

 

 

Fallow Deer

£22.95

 

 

 

Muntjac and Water Deer: Natural History, Environmental Impact and Management

£34.99 

All prices correct at the time of this article’s publication.

NHBS In the Field – Elekon Batlogger M2

The Elekon Batlogger M2 is a sophisticated bat detector, designed for use in active transect surveys and bat walks. The detector is ergonomically designed and compact in its build. Unlike the Batlogger M, the replaceable microphone is set back within the casing, reducing the chance of breakages if accidentally dropped.

It is an easy-to-use detector, able to record ultrasonic calls over a wide range of 10-192Hz, retaining details of the harmonic structure and amplitude of the original bat call.

This detector also includes features for real-time heterodyne monitoring scheduled recording and several preset recording modes for typical survey types, including ‘Explore’, ‘Transect’ and ‘Passive’. There is also an option to create your own presets, with all settings accessible directly on the Batlogger M2.

The recordings can be easily and quickly managed and analysed using the free BatExplorer Software for Windows. This software also includes computer-aided species identification.

How we tested

In August 2021, we tested the Batlogger M2 on Dartmoor in South Devon at dusk. We chose an area with both open moorland and woodland to monitor the dusk emergence of several bat species. Using the ‘Explore’ recording mode, the device was set to real-time heterodyne monitoring, with the trigger settings selected to 45Hz.

The recordings were stored on a 16GB MicroSD card, output via WAV and XML. The files can be easily transferred to your computer with the included cables (a USB-C to USB-C cable and a USB-C to USB-A adapter) however, we simply used an SD card reader to access and transfer our recordings.

Elekon’s ‘BatExplorer Lite’ software was used to visualise and analyse our recordings.

What we found

At only 222g, this device is lightweight, therefore it was not a hindrance when hiking to the site. It was easy to hold and operate in low light, which is important as it will often be used in conjunction with other equipment while undertaking surveys. However, the buttons do not light up or glow in the dark, therefore it is important to familiarise yourself with the setup beforehand.

Our survey used the ‘Explore’ recording mode however, we looked briefly at the other preset modes and felt they would be very useful if carrying out these types of surveys. In particular, the ‘Transect’ mode allows you to pre-program the detector with your own GPS survey route, which will then display directions to keep you on path.

Setting up our recording session was quick and easy. When triggered, the detector automatically recorded the calls in full-spectrum, therefore no further operation was required. Each recording logs the GPS location, temperature, humidity and brightness at the time of recording, so there is no need to bring separate equipment for these parameters. The device also has an integrated voice microphone to enable you to take time-stamped voice notes while surveying, which allows for more accurate note-taking than writing in the dark.

The screen display was bright and easy to view in the dark. The main screen shows the current recording time and frequency, as well as amplitude and harmonic structure of the previous bat call, the time since the last sequence and the species suggestion for that recording. The other recorded information can be accessed by pressing the left and right buttons to switch screen displays, allowing for an uncluttered screen and fewer distractions.

We tested the detector’s audio output with both earphones and through the built-in speakers (includes an adjustable volume). Both had clear audio with little background noise.

We recorded multiple calls including over 60 calls during one 30 minute survey. We were impressed with the quality of the recordings and the lack of ambient noise assisted in the identification of calls.  It is worth noting that the M2 uses a SiSonic microphone, whereas the older Batlogger M uses an FG Knowles microphone, so experienced Batlogger M users may notice a slight difference in their recordings.

Automatic species identification was not always to species level, with some recordings having no suggested species. Those that were suggested, however, appeared mostly accurate when we analysed the recordings via the BatExplorer software.

Our opinion

The Batlogger M2 by Elekon is a very easy-to-use and accessible detector. The preset recording modes are useful for almost all circumstances and allow quick survey set up without the need to trawl through complicated settings – particularly helpful for newer ecologists. The only limitation we found was the lack of visual aids for the buttons in the dark, however, this can be easily overcome by familiarisation and repeated use.

It is clear from the design of the M2 that Elekon has acted on feedback from the Batlogger M and have created an intuitive detector with a robust build, perfect for any survey needs.


The Elekon Batlogger M2 can be found here. Our full range of active bat detectors can be found here.

If you have any questions about our range or would like some advice on the right product for you then please contact us via email at customer.services@nhbs.com or phone on 01803 865913.

 

The NHBS Guide to UK Rabbit and Hare Identification

Rabbits and hares are species in the family Leporidae, which contains over 60 species. They are mammals within the order Lagomorpha, together with the pikas. They are small to moderate-sized species, characterised by long hind legs, long ears and rapid movements. They are almost exclusively herbivorous, feeding mainly on grasses and herbs, although they do also eat leaves, fruits and seeds. Leporids are coprophagous, meaning they pass food through their digestive system twice. To do this, they first expel the food as soft green faeces, termed cecotropes, which they then reingest, eventually expelling it again as dark faecal pellets. This increases their ability to break down and digest plant material, extracting further nutrients.

They inhabit a wide range of habitats, from mountains and wetlands to forests and grasslands. Leporids play an essential role in many of these ecosystems, as seed dispersers, ecosystem engineers (a species that significantly modifies their environment) and as a primary prey item for a number of predator species, such as foxes, wildcats and some mustelids, including stoats and weasels. Their young are particularly vulnerable to these predators, as well as badgers, domestic cats and several birds of prey. Because of this vulnerability, many leporids have large litters, often nesting in burrows underground to protect their young from predation.

One threat to populations, particularly in European rabbits, is myxomatosis, a disease caused by the poxvirus Myxoma virus. Its two natural hosts, the tapeti and brush rabbits of South, Central and North America, experience only mild disease. However, myxomatosis is a severe and usually fatal disease in European rabbits. When the disease originally spread to the UK in the early 1950s, the mortality rate was 99%. In the 1970s this declined to between 47 and 69% but populations were severely affected. The disease, which causes localised swelling, skin lesions, blindness and respiratory distress, has also been deliberately introduced into the wild on multiple occasions. Used to try to eradicate or control rabbit pest populations, myxomatosis was intentionally introduced in Australia, New Zealand, South America and parts of Europe, including Britain.

Other threats to leporid populations include rabbit haemorrhagic disease (RHDV1 and RHDV2), hunting, habitat loss and agricultural intensification. Rabbit populations declined by 64% in the UK between 1996 and 2018 and numbers of brown, mountain and Irish hares are also thought to have declined in some areas. As ecosystem engineers, the loss or reduction of these species can have major consequences, particularly for rabbit-dependent habitats. In their absence, the consequent changes in vegetation structure due to a lack of grazing can have further impacts on other wildlife, such as invertebrates.

Rabbit (Oryctolagus cuniculus)

Distribution: Widespread across the UK, although they are absent from Rum, Isles of Scilly and some smaller islands.
What to look for: Rabbits are the smallest leporids in the UK, with the characteristic long ears and long hind legs. They have a sandy, grey-brown colouration and a white tail. They have dark, amber coloured eyes. Other than their size, the most noticeable difference between rabbits and other leporids in the UK is the lack of a black tip on the ears.
Did you know? The rabbit, also known as a coney, is not native to the UK. The exact date of their introduction is one of ongoing research. It was previously thought that rabbits were first introduced by the Normans in the 11th or 12th century as both a food and fur resource. But the recent re-examination and radiocarbon dating of a bone found at a Roman palace show that at least one rabbit was present much earlier, in the first century AD. Researchers have stated that there is no evidence of many rabbits in the area and another analysis suggests the rabbit was kept in confinement, therefore it has been suggested that it was most likely kept as an exotic pet.

JJ Harrison via Flickr
Brown hare (Lepus eueopaeus)

Distribution: Widespread across the UK, although they are less common and more restricted in Northern Ireland and absent from parts of north-west Scotland.
What to look for: This species is larger than the rabbit, with long, black-tipped ears and very long hind legs. Their colouration is redder than the rabbit and the mountain hare, and they have a black-topped tail that is white underneath. The brown hare has amber eyes and their fur can appear grizzled.
Did you know? This species is also non-native and were introduced in the Iron Age. Radiocarbon dating of bones found in Hampshire and Hertfordshire, along with historical accounts, suggest that brown hare were not eaten until hundreds of years later during the Roman period, and were instead associated with deities.

caroline legg via Flickr
Mountain hare (Lepus timidus)

Distribution: Mainly in the highlands of Scotland, although they are also found in other areas of Scotland, on some Scottish islands and in the Peak District.
What to look for: This species has a grey-brown coat during the summer and a white coat in the winter, although the tips of their ears remain black in both forms and their tail remains white. They have brown eyes and long ears, although these are shorter than the brown hare’s.

Brown form: Andrew via Flickr
White form: John Johnston via Flickr
Irish hare (Lepus timidus hibernicus)

Distribution: Widespread across Ireland
What to look for: They are very visually similar to the mountain hare, except they do not develop a white coat during winter months and are noticeably smaller. During the summer, their coat is reddish-brown which dulls to grey-brown in winter, although their tail remains pure white.
Did you know? The Irish hare is a subspecies of the mountain hare and is the only lagomorph species native to Ireland. Fossil carbon dating suggests that these hares have been present in Ireland for at least 30,000 years.

Jimmy Edmonds via Flickr (Image cropped)
Domestic rabbit (feral) (Oryctolagus cuniculus domesticus)

Distribution: Unknown
What to look for: Domestic rabbits can have a huge variety of appearances, with at least 305 breeds of domestic rabbits around the world, although different numbers are accepted by different organisations. They can vary in size, colouration, body shape, coat type and ear length.
Did you know? Many pet rabbits are abandoned each year, particularly in the period after Easter.

Paul Korecky via Flickr
Suggested reading:

Britain’s Mammals: A Field Guide to the Mammals of Great Britain and Ireland
£17.99

 

 

 

RSPB Spotlight: Hares
£9.99

 

 

 

Guide to the Land Mammals of Britain
£3.75

 

 

 

How to Find and Identify Mammals [Revised Edition]
£11.99

 

 

 

All prices correct at the time of this article’s publication.

Gift Ideas That Support Wildlife

This festive season, why not consider giving a gift that will also support your local wildlife. Wildlife populations in the UK are facing serious threats and many species are in decline, however there are ways in which we can protect and help at-risk species by creating havens for wildlife in our own gardens. At NHBS we sell a range of products, from bird feeders to hedgehog houses, that can both bring joy to the recipient and benefit wildlife at the same time. We also sell a number of books that can help you create a wildlife friendly garden. We’ve put together a selection of some of our favourite items for you to browse below. 


Eco Hedgehog Hole Fence Plate

#242607 

Hedgehog numbers have dramatically declined in recent years. Creating a hole in a garden wall or fence will allow your local hedgehogs to pass through from garden to garden safely.

 

NHBS Wooden Bird Nest Box

#254495

Many bird species are struggling to find enough suitable natural nesting sites in the modern environment, but a bird box will provide a warm, sheltered substitute, with protection from most types of predators, helping to improve the chances of breeding success.

 

Froglio Frog and Toad House

#216744

The Frogilo Frog and Toad House provides a safe retreat for frogs and toads in any garden and is handmade in frost-resistant ceramic with a decorative glazed roof.

 

National Trust Apex Insect House

#251682

The National Trust Apex Insect House is an ideal addition to any wildlife friendly garden. With a variety of shelter types, it offers a perfect habitat for important invertebrates such as lacewings, ladybirds, and even some butterflies.

 

Bee Brick

#244140

Bee Bricks are made in Cornwall in England using the waste material from the Cornish China clay industry.  They provide much needed nesting space for solitary bee species such as red mason bees and leafcutter bees, both of which are non-aggressive.

 

Echoes Bird Bath

#195520

A large and beautifully coloured and glazed bird bath with a ‘ripple’ step design that is both visually attractive and functional by providing extra footing/grip for wild birds.

 

Defender Metal Seed Feeder

#238813

The Defender Feeder’s metal construction is tough, long lasting and offers excellent protection from squirrel damage.  The feeder is available with two, four or six feeding ports, each with a perching ring that allows birds to feed in a natural, forward facing position.

 

 

Hedgehog House

#234035

Hedgehog numbers are rapidly declining across the UK and providing a refuge in your garden with the Wildlife World Hedgehog House will help to protect hedgehogs from predators and disturbance.

 

RHS The Little Book of Wild Gardening

#257312

This is a guide for anyone wanting to garden in a more sustainable, natural way. Working with nature benefits not just the garden, but also the gardener, wildlife and the wider environment.

 

Gardening for Bumblebees

#252488

This shows you how you can provide a refuge for bumblebees to feed, breed and thrive. No matter how large or small your space is, Dave Goulson shows you how you can make a pollinator-friendly haven.

 

 

The Wildlife Pond Book

#246688

This offers a fresh and unique perspective on ponds, encouraging readers of any budget to reach for the spade and do something positive to benefit their shared neighbourhood nature.

 

Wildlife Gardening

#244291

If you want to attract more bees, birds, frogs and hedgehogs into your garden, look no further than this. Kate Bradbury offers tips on feeding your local wildlife and explains how you can create the perfect habitats for species you’d like to welcome into your garden.


Discover more great gift ideas on our website. Plus, check out our two blogs on how to attract wildlife to your garden.