ID guide – Hoopoe – A blog by nhbs

Author interview with Steven J Falk: A Photographic Guide to Flies of Britain & Ireland

Over recent decades various ‘difficult’ insect groups have gradually been brought within reach of the non-specialist thanks to developments in field guides. Flies, however, have remained something of a final frontier, and it’s easy to see why – with more than 7,000 species in Britain alone, creating an accessible and user-friendly guide to this group is a daunting task.

Now, however, a team of expert field entomologists have stepped up to the challenge and produced A Photographic Guide to Flies of Britain and Ireland – the first guide to ever tackle this amazingly diverse insect order as a whole. With beautiful photography and clean, concise text, this book is set to put flies on par with bees and other more ‘popular’ insect relatives, and bring them their due share of attention in the interests of general naturalists.

Ahead of publication, we had the pleasure of talking to lead author Steven Falk about the book’s development and the rewards of fly-recording.

Firstly, how did the book come about?

The idea for the book initially came from Peter Creed, the Creative Director (and Designer) at NatureBureau, which owns Pisces Publications. He wanted a book resembling A comprehensive guide to Insects of Britain & Ireland by Paul Brock, published by Pisces in 2014 (with a 2019 second edition) but concentrating purely on Diptera. He approached Gail Ashton who was helping to man the Dipterists Forum stand at one of the Amateur Entomology Society autumn fairs to discuss the possibility of doing such as book in conjunction with the Forum. Gail eventually agreed to help write it and asked Rory Dimond to assist. I become involved about one-third of the way through the project when it was clear that the book would need several hundred of my images to make it viable (we needed at least 1,500 accurately identified fly photos for the book), and my role later expanded into that of primary author. The internet has patchy information on Diptera, and what does exist can be misleading or out of context with the British fauna, so you need lots of first-hand experience to avoid creating misleading text. Even taking information from trusted literature can be challenging when it needs it be distilled down to short, snappy species accounts. Gail and Rory helped ensure that the text was not overly technical, though inevitably a book on flies ends up more technically worded and microscope-based than a book on butterflies or dragonflies. But we provide good diagrams and a large Glossary to help readers negotiate this. The proof-reading and creative discussions by the members of the team have been very important, making for a much better publication. It helps that all four of us share a passion for breaking down barriers in entomology and enthusing general naturalists about flies rather than just serving the converted. We were also keen to produce a more visually exciting and enticing book than much of the more technical fly literature and online resources available for flies. But we have still signposted those other resources for readers who want to develop a deeper understanding of flies and take things further by providing a good Bibliography. This covers all the key identification literature relevant to a British or Irish audience, both printed and that which is freely downloadable from the internet.

Let’s talk about flies themselves. What makes them so important, and what can recording flies tell us about the state of the natural world?

There are currently about 7,300 species of fly on the Britain and Ireland list, with the possibility of over 10,000 species being present, as we find dozens of new ones each year and DNA is revealing lots of cryptic species. So, if you are passionate about biodiversity and serious about its conservation, you cannot ignore flies. They are typically the most speciose eukaryotic group at a wildlife site and are also incredibly diverse ecologically. I regularly record over 500 fly species at a good wildlife site, and the Windsor Forest and Great Park list stands at almost 2,000 species. Those massive fly assemblages including predators, parasites, saprophages, herbivores, fungivores, pollinators, and so on. What is more, fly assemblages can be found from the intertidal zones of the coast to the tops of our highest mountains. Having some idea of what all those flies are and what they need in terms of breeding sites and adult feeding, makes you view habitats, sites and landscapes in a very different way to a botanist or general ecologist. You notice and value microhabitats and other features that others miss or under-value, such as sap runs or water-filled rot holes in old trees, different sorts of dung, different sorts of ‘wet’, different sorts of ‘decay’, different sorts of ‘bare ground’ and so on. Flies also make you think much harder about aspects such as habitat combinations and mosaics (bearing in mind that fly larvae and adults often have very different needs), habitat connectivity, habitat condition, site history, microclimate, seasonality and climate change. Dipterists are often pretty good botanists and habitat ecologists, but adding flies and other insects to the equation provides a much bigger ‘vocabulary’ when you are trying to understand and interpret the environment. That was particularly important a few years ago when several professional entomologists including myself were assessing the emerging Biodiversity Net Gain metrics. We were able to articulate some powerful feedback to Defra and Natural England concerning the over-simplistic metrics that were emerging, and the serious impact that might have for protecting invertebrate (as well as general) biodiversity. Flies are also really important in ecosystem services (e.g. crop pollination, pest control, sewage treatment). There is strong evidence that they may be more important than bees for the pollination of certain crops, and for general wildflower pollination in habitats like montane grassland and saltmarsh. The role of flies in ecosystems and food webs is something that comes through strongly in the book.

Faced with such a huge group, how did you go about deciding which species to include?

Being a photo-based book, the choice of what species to include is heavily based on what photos are available. But many fly photos on the internet are inaccurately identified, so we had to make sure that such images were either avoided, or (if we could tell what they actually were), used correctly. So, if a species lacked a decent photograph, it could not be subject of a typical species account, though we have name-checked many species without accompanying images within the Similar Species section at the end of many species accounts. For some smaller, iconic families, such as robber flies, horseflies, soldierflies and bee-flies, every published British and Irish species is included. For hoverflies, it is closer to about two-thirds of the fauna, and for groups like parasite flies and the housefly family, it is about half the fauna. In these instances, we tried to include representatives from most if not all the genera in a family and tried to ensure most of the more distinctive-looking or ecologically interesting species were covered. For some of the more obscure or difficult families, only brief coverage is provided. But all 108 fly families present in Britain and Ireland get some level of coverage in the book, which is quite an achievement.

Can you tell us a bit about the book’s approach – what can readers expect from the species accounts?

The book is arranged taxonomically following the sequence of the 2025 Checklist of Diptera of the British Isles which is hosted (and regularly updated) on the Dipterists Forum website. The individual species accounts mostly have a consistent format. It starts with an indication of the size of a fly, either using wing length or body length, then a basic description of the species, often carefully worded to highlight differences from other similar species. We then describe Habitat, Distribution, Season, and (for parasitic or herbivorous species) Hosts. The species accounts are accompanied by a map and photograph or, indeed, two or more photos for species with strong sexual dimorphism or several colour forms. A section on Similar Species is also used in many cases, namechecking other flies that might be confused with the main subject. These extra species are also sometimes provided with a map and photo. It means that about 1,300 species of fly get namechecked by the book (about 1,100 of these with standard species accounts). But it often came down to what space was available on each page spread as to what got covered. Suffice to say we used a flexible and opportunistic approach to ensure we made the most of each double-page spread.

The guide is beautifully laid out and packed with outstanding imagery. Was it a challenge gathering photos for all the flies you wanted to feature?

About 40% of the photos come from me and were already featured on my Flickr site. That made image selection for some families much easier and meant that Peter Creed could quickly download any of my images, and if I disagreed with an image choice, I could quickly send him hyperlinks to the image I preferred. Peter, who is also a keen insect photographer (and always thinking of the next potential Pisces book), provided about 20% of the images. We also turned to reliable British insect photographers such as Paul Brock, Simon Knott, Kevin McGee and Ian Andrews, and approached other photographers if we spotted images of further species on the internet (or better images of a species than we had to hand). The result is that we’ve ended up with over 1,500 images of over 1,100 species and have used images from 186 photographers. Suffice to say, we are immensely grateful to everyone who provided images.

For the general naturalist who has yet to become a fly convert, what can you say about the rewards of studying these insects?

Flies have huge intrinsic interest in terms of interesting appearances (including mimicry, loss of wings, bizarre wing markings and strange body modifications), interesting lifecycles, interesting behaviours, and their importance in ecosystem services. They are great fun to photograph (with cameras like the Olympus TG series making it easier than ever), and it is relatively easy to identify some from photos, either using this book, or by posting images on the various Facebook groups that cover flies (where others can provide feedback on what they think your photo is). Flies also provide an incredibly powerful framework or lens for interpreting and understanding habitats and landscapes better, as explained above. I think that aspect (using flies to view understand the environment more critically) is truly exciting and very rewarding.

What tips would you give for someone looking to take a deeper interest in flies and find a greater range of species?

Buying a stereo zoom microscope makes a big difference as it allows you to identify flies with much greater confidence and appreciate the beauty in the details of their morphology. While we are aware that many users of the book will not want to go down the road of collecting and killing flies for the purpose of identifying them critically, that is the approach that is generally needed to develop long and accurate species lists for a site. There is a limit to what can be achieved through photography. But going down the collecting route also means buying an insect net, tubes and pooters, insect pins and storage boxes for pinned specimens, plus obtaining some of the key literature listed in the book’s extensive bibliography, such as British Hoverflies (Stubbs & Falk 2002), published by the British Entomological and Natural History Society. I would also strongly recommend joining the Dipterists Forum. This is the national society for the study of flies and is very friendly and well-organised. It publishes a regular newsletter (Bulletin of the Dipterists Forum) and journal (Dipterists Digest) and organises regular indoor and outdoor events. It has a great website that can be used by non-members but has extra resources for logged-in members. The Forum acts as an umbrella for almost 30 family recording schemes or study groups, some of which have their own Facebook groups or satellite websites. Nowhere outside of Britain and Ireland comes to close to matching that offer.

Finally, what do you hope the book will do for interest in flies and fly recording?

We are hoping it has a major impact, not only by making general naturalists more aware of, and sympathetic towards flies but also by encouraging more naturalists to take up formal recording of flies as outlined in the previous response. Many parts of Britain have few (if any) resident fly recorders, and it would be especially good to promote recording in those areas, even if it is just the recording of easier groups such as hoverflies. Climate change is having a profound impact on insect life, and the more data we get, the more we understand that impact and formulate strategies to counter it through improved habitat management and other land use decisions. We hope that this book will support constructive dialogue – ecologists, farmers, planners all a little bit more familiar with flies, able to be enthusiastic about them and their conservation.

A Spotlight on the Deadly Mushrooms of Britain

The UK boasts a rich and diverse fungal kingdom, from spongy puffballs to otherworldly slime moulds and foul-smelling stinkhorns – among these edible treasures lurk several deadly species, capable of causing serious illness and even death. In this blog, we will explore some of the most notorious poisonous mushrooms in the UK.

This blog is for educational purposes only and should not be considered a definitive guide to mushroom identification. Always consult with a professional before consuming any wild mushrooms.

Fly Agaric (Amanita muscaria), also known as Fly Amanita

Commonly depicted in fairy tales, artwork and film, Fly Agaric is a familiar image that comes to mind when one thinks of fungi.

Identification: These common mushrooms are most readily identified by their contrasting bright red (or in some cases, orange) cap and white flesh. The cap can reach 20cm across, is initially hemispherical in shape before opening flat as the mushroom matures and is often accompanied by white spots or ‘warts’ that can be washed off with rain – a remnant of the ‘egg’ covering. On the underside, the gills are closely packed together and are not joined at the stipe – a brittle, bright white stalk. The stipe features a large white skirt that may have yellowed edges, and at its base is a bulbous volva (a cup-like structure) covered with shaggy, scaled rings. Spores are oval and white. Fly Agaric are widely distributed across heathland and moorland from August to December, and show a preference for light soils among birch, pine or spruce trees with which they form mycorrhizal associations.

Toxicity: Reported symptoms of Fly Agaric poisoning can vary greatly, but primarily, this mushroom causes hallucinations, psychosis, diarrhoea and digestive upset. It can be mistaken for the Blusher (Amanita rubescens), an Amanita which has a similar shape, and a pale red-brown cap with cream spots.

Fungi fact: This species was traditionally used as an insecticide – where the name Fly Agaric originated. Ibotenic acid within the toadstool attracts flies and subsequently kills them, which led to its use as a medieval fly trap, where it would be broken up and sprinkled into milk. It has also been used in Siberian civilisations, where it has been consumed for religious and recreational purposes for millennia.

Panthercap (Amanita pantherina)

The Panthercap is another distinctive member of the Amanita family, and is strikingly similar to its vibrant cousin, Fly Agaric.

Identification: This poisonous toadstool is not particularly common in Britain, but can be found from July to November in broadleaved woodland, where it forms ectomycorrhizal relationships with beech and oak trees. It can grow up to 14cm in height, with a cap width of around 12cm for mature specimens. The cap itself flattens with age, appears dark brown to slightly red in colour, and is usually covered with thick white ‘warts’ or spots. Underneath the cap, the crowded white gills are free from the stipe and produce white spores that are broadly ovate in shape. The white stipe typically appears smoother towards the top and shaggier under the skirt, which is generally smooth and white. The flesh inside the stipe and cap is hollow and white.

Toxicity: Panthercaps toxins can result in suppression of the central nervous system, involuntary muscle movement, hallucinations, gastrointestinal upset and an increased heart rate. These toadstools may be mistaken for the Blusher, as well as the Grey Spotted Amanita (Amanita excelsa) which can be distinguished by striations on the upper surface of the skirt.

Funeral Bell (Galerina marginata)

Funeral bell mushroom — Image by Andre De Kesel via Flickr

A highly toxic little brown mushroom, the Funeral Bell can quite easily be mistaken for other edible species. It can be found fruiting in clusters on woodchips, dead rotted wood and coniferous/deciduous tree stumps from August to November.

Identification: Funeral Bells can grow up to 7cm in height, with a cap diameter of approximately 6cm. The cap can be pale yellow or brown in colour, or orange in some cases, and the edges are usually lighter than the centre. Pale tan to red/brown gills on the underside of the cap are relatively crowded and are broadly attached to the stipe, occasionally running down it slightly. The fibrous stipe can be found in varying colours, from pale tan to dark brown, which depends largely on maturity, and it has a small membranous skirt, with pale, thin flesh that darkens as the fruit matures. Spores are brown in colour.

Toxicity: Funeral Bells have the same toxins as Death Caps and Destroying Angels – when ingested, these compounds cause gastrointestinal upset within 24 hours, before suddenly resolving. Symptoms return shortly after this and quickly progress into organ failure and eventual death. The Funeral Bell can be easily mistaken for other little brown mushrooms and has been mistaken for Velvet Shank in the past – easily distinguished by a black velvety stem and the absence of a skirt.

Fungi fact: One single mushroom contains enough toxin to kill an adult human.

Deadly Webcap (Cortinarius rubellus)

Although rare in the UK, Deadly Webcaps can be found in coniferous pine and spruce woodland, growing among heather or bilberry from August to November.

Identification: On average, these mushrooms grow to 8cm in height, with a cap width of up to 6cm. Young caps start with a convex shape, before maturing into an umbonate form with a raised and slightly pointed centre. The cap colour can range from reddish brown to orange and the surface may appear hairy or scaly. On the underside, gills can be orange, brown or red, often appearing paler on immature toadstools, and spore prints of this species are typically rusty brown. The gills themselves are fairly widely spaced and emarginate, joined to the stipe with a slight notch and younger specimens may have a pale, slightly orange, cortina (or veil with a cobweb-like structure) that cover them. The stipe can vary in colour and has a slightly bulbous lower half, tapering towards the base.

Toxicity: Following ingestion, symptoms can take up to two weeks to develop, manifesting as digestive upset and interruptions of the central nervous system, soon followed by kidney and liver failure.

Fungi fact: Deadly Webcaps are reported to have a scent reminiscent of radish.

Death Cap (Amanita phalloides)

Death Caps are fairly common in broadleaved and mixed deciduous woodland with oak and beech trees, where they can be found in widely spaced groups from July to November.

Identification: On average, these mushrooms grow to around 15cm in height. They begin with a convex cap that flattens as it matures and can be olive yellow to green-bronze in colour – depending on the age of the fruit, a white cap may also be observed. The cap is often darker in the middle and has radiating fibres that give a streaked appearance. Underneath, the gills are broadly spaced and are free of the stipe, appearing white, cream or pink in colour, which changes as the fungus matures. The stalk is generally off-white in colour with a clearly striated skirt and an obvious deep volva. Death Cap spores are white and elliptical in shape.

Toxicity: Death Cap poisoning starts with gastrointestinal symptoms several hours after ingestion. These symptoms can last for days before a brief recovery, after which the liver and kidneys fail resulting in death in most cases. This deadly mushroom can easily be mistaken for a number of edible varieties but can easily be distinguished by a sickly-sweet smell.

Fungi fact: Death Caps have caused the most fatalities from mushroom poisoning in the UK and are known as the world’s most deadly fungus. It has been used as a murder weapon for millennia, and there is no known antidote to its toxins.

Destroying Angel (Amanita virosa)

Destroying Angels are relatively uncommon in the UK but can be found in broadleaved and mixed deciduous woodland, especially those with Birch trees, from July to November.

Identification: Destroying Angels stand up to 14cm in height, with a cap width of 11cm. They are pure white in colour and start with a convex cap, before maturing to a conical shape and eventually lying flat. Underneath, the white gills are crowded and free of the stem, which has fibrous skin and a bulbous volva at the base. Destroying Angels have a skirt on the upper half of the stipe, which is fragile and may be missing or small in size – it is not striated and is also white in colour. The flesh inside the mushroom is pure white, and the spores are white globose. It is has an unpleasant sickly-sweet smell.

Toxicity: Symptoms – vomiting, diarrhoea and abdominal pain – begin several hours after ingestion and can last for several days before easing into a false recovery and eventual organ failure. Destroying Angels may be mistaken for edible Wood Mushrooms (Agaricus silbicola), which can be distinguished by the smell of aniseed and brown spores.

Fungi fact: Small mammals, including rabbits and squirrels, are known to consume Destroying Angels with no ill effect.

The NHBS Guide to UK Wasps

Often seen as mere picnic nuisances, the UK is home to a surprising variety of wasps, including both social and solitary species. While many of us will be familiar with the iconic black and yellow social wasps, numerous solitary species display incredible diversity in both form and behaviour. Distinguishing them can seem daunting, but key features like body shape, coloration, and reproductive behaviour can offer valuable clues.

Here, we’ll explore a selection of common wasp species, and some wasp families found across the UK, detailing their identifying characteristics and the typical habitats they can be found in.

**Common European Yellowjacket (*Vespa vulgaris***)

Common wasp DSC_5493_edited-1 by Clifton Beard, via flickr.

Identification: Between 11-19mm in length with striking black and yellow stripes. These social wasps have six abdominal sections, each with one stripe, and there is a very distinguished waist between the thorax and the abdomen, which has black dots and rings. The species has a conspicuous face on a triangular shaped head with a black anchor or dagger shaped mark towards the base, and a pair of long, robust antennae. They have yellow cheeks and yellow legs.

Distribution: Widespread throughout the UK. Common Social Wasps can be found in almost all habitats, particularly grassland, heathland and woodland from mid-March to November.

**Red Wasp (*Vespula rufa***)

Rote Wespe (Vespula rufa) by Frank Vassen, via flickr.

Identification: Growing up to 20mm in length, Red Wasps can easily be distinguished from V. vulgaris by reddish-brown markings on their upper back segments.

Distribution: Widespread, and frequently found in open woodland, moorland and hedge banks from the end of March until September.

**Hornet (*Vespa crabro***)

Identification: Hornets are Britain’s largest social wasp species. Around 15-35mm in size. The abdomen is yellow and unbanded, with brown markings on the upper parts, while the thorax is black and reddish-brown. Their legs are a reddish brown – a characteristic that can distinguish them from Asian Hornets – and antennae are yellow or brown in colour.

Distribution: Widespread distribution and commonly found throughout England and Wales. Hornets can be found in many lowland habitats and are known to associate with ancient woodland, wood pasture and can be found in parks and gardens in more urban areas. Queens emerge from overwintering in early April, while workers can typically be seen from late June to November.

**Yellow-legged Asian Hornet (*Vespa velutina***)

Asian hornet by Gilles San Martin, via flickr.

Identification: Typically smaller than native hornets, this invasive species can grow up to 3cm in length. Their abdomen is dark in colour with fine yellow stripes, and the fourth segment towards the base of the hornet is distinctly yellow orange in colour, while the thorax is entirely black or dark brown and appears velvety in texture. From the front, the head appears orange and the legs have yellow tips.

Distribution: Yellow-legged Asian Hornets were first recorded in England around 2016, after they were accidentally introduced from Asia. They have mostly been recorded in the south of England, particularly the South-east.

Wasp Families

The diversity of wasp families in the UK is vast and presents a significant identification challenge. In this blog, we have chosen to focus on a number of wasp families to demonstrate the wonderful diversity of this group.

Ichneumon Wasps

Sabre Wasp – Rhyssa persuasoria by Nature Spot. Ichneumon wasp by nutmeg66, via flickr. Barylypa propugnator by Nature Spot.

There are around 2,500 species of Ichneumon wasp in the UK. These small insects can vary in size but generally reach up to 1 ½ inches in length. They exhibit a wide range of colours and patterns, some with brightly coloured bands, and others with more subtle markings. Ichneumon Wasps can be distinguished by a slender sickle-shaped abdomen and a pair of long antennae, each with at least 16 segments. They have a narrow waist, long jointed legs and two pairs of wings. These parasitic wasps inhabit flower-rich meadows, gardens and woodland – they can often be seen around umbellifers or investigating vegetation and dead wood.

Gall Wasps

Oak Marble Gall by Nick Wood, via flickr.

The UK is estimated to have 86 Gall Wasps, a group of well-studied parasitic wasps. This group gets its name for a unique reproductive behaviour where adults lay eggs in the soft tissues of trees, flowers and other plants, creating a gall as the larva develop. Usually on the underside of leaves, branches and around acorns, these wasps produce conspicuous galls on their host plant – interestingly, they usually only infect a specific species of genus, for example, Citrus Gall Wasps will exclusively parasitise citrus trees. After overwintering, the larvae will emerge in spring and can be seen throughout summer. They are typically under 5mm in size, shiny, and are black or brown in colour with a humped thorax and wings.

Cuckoo Wasps

Crysis cf impressa close-cropped by Nigel Jones, via flickr. Chrysis fulgida by Ryszard, via flickr. Chrysis viridula by Tim Worfolk, via flickr.

Instantly recognisable by their brilliant metallic body colours in shades of blue, green, purple and red, these small chrysidids are also known as jewel or ruby tailed wasps. There are around 31 species of Cuckoo Wasp in the UK, which can be seen in sunny weather from April to October in a range of habitats; particularly in sandy soils around coastal areas, gardens and ancient woodlands. The group are often seen investigating the nests of other Hymenoptera species in order to reproduce – these parasitic species will enter the host nest and lay their own eggs in the host cells. Once hatched, Cuckoo wasp larvae will eat the eggs and larvae of the host species before restarting the cycle – hence the name ‘cuckoo’.

The NHBS Guide to UK Orchid Identification

Orchidaceae, a family boasting over 28,000 species worldwide, are renowned for their intricate beauty and fascinating adaptations. These perennial plants exhibit a remarkable diversity of forms, even within species, with flowers featuring three sepals and three petals, one of which often forms a distinctive lower lip (the labellum) which is an important identification feature. They rely on diverse pollination methods, often involving intricate relationships with insects and fungi, and are highly specialised to survive in specific habitats.

The UK is home to 57 native orchid species, ranging from common meadow flowers to prized rarities. This blog explores a group of these British orchids and provides information on both their identifying characteristics and distribution in the UK.

Bird’s-nest Orchid (Neottia nidus-avis)

birds nest orchid among grass — Image by Bjorn S via Flickr

Identification: Bird’s-nest Orchids are incredibly distinctive and are named for their unique root system that resembles a bird’s nest. They have no leaves, and do not have chlorophyll, so they rely on nutrients provided by mycorrhizal fungi in the soil.

The cylindrical spike grows up to 50cm in height and can carry up to 100 yellow-brown flowers that smell like honey. The sepals and upper petals are short and form a fan-shaped hood at the top of the flower. At the base is a broad, long lobe that is forked in the middle, and at the base of this lip is a shallow cup of nectar which is used to attract small insects for fertilisation.

Distribution: This species has a scattered distribution throughout the UK but is locally common in southern England and Northern Ireland. Bird’s-nest Orchids can be found in mature woodland, especially those with beech and Yew, in southern England and typically flower from early May to late June.

Burnt Orchid (Neotinea ustulata)

Identification: This distinctive orchid has two variations: early flowering and late flowering forms, which exhibit subtle differences in flower colour. This orchid is conical to cylindrical in shape and features densely packed, deep-purple flower buds. These flowers have dark purple-red sepals and pale petals, including a four lobed white lip with red-purple spots.

Early Flowering Form: Grows up to 10cm tall, with a closely packed flower spike bearing up to 50 small flowers. The sepals and upper petals form a tight, darkly coloured hood, which appears red-brown when the flowers first open and quickly fades as the bud matures. It has a long white lip with two rounded side lobes, and a bluntly forked central lobe. Early flowering forms have a strong, sweet scent and are pollinated by flies.

Late Flowering Form: Typically found between 8 and 15cm in height. Late flowering form flowers have shorter, blunter lobes and the red colouring on the hood remains strong even with maturity. The edges of the lip are usually flushed with colour, which can spread over the whole lip in some cases.

Distribution: Formerly widespread throughout the UK but has undergone declines in the north and midlands. It is now found in a few localities, mainly in southern England, on chalk and limestone grassland and meadows. Early flowering forms bloom mid May to mid June. Later flowering forms bloom late June to early August.

Greater Butterfly Orchid (Plantanthera chlorantha)

Identification: Greater Butterfly Orchids can grow up to 60cm in height. The plant consists of a tall spike of up to 40 greenish-white flowers in a loose cluster. The overall flower structure is similar to Lesser Butterfly Orchids and has ovate lateral sepals with a wavy edge like the wings of a butterfly. The upper sepal and petals form a broad, green-white hood and a long, translucent lip has a green tip. At the base of the stem, there are two large, shiny leaves that are blue green in colour and elliptical in shape.

Distribution: Widespread throughout the UK, but most common in southern England. Found in meadows, open scrub, grassland and ancient woodland on chalk grassland or calcareous soil. Flowers from late May to late July, where they emit a powerful scent at night and are pollinated by moths.

Common-spotted Orchid (Dactylorhiza fuchsii)

Identification: Up to 70cm in height, the spike is either pyramidal or cylindrical in shape and can host up to 70 flowers at a time, all various shades of pink, purple and on occasion, white.

The flowers are slightly scented and are pollinated by a range of insects but are particularly attractive to day-flying moths. They have a three lobed lip with a larger, more triangular central lobe that is marked with a pattern of dark pink spots and stripes. The lateral sepals are spreading, and dorsal sepals and upper petals form a loose hood at the top of the flower. All petals and sepals are marked with dark pink lines and dots. It has spotted, narrow basal leaves that form a rosette at ground level.

Distribution: One of the most common and widespread orchids in the UK, but is absent from much of Cornwall, Devon and northern Scotland. They can be found in a range of habitats, including woodland, dry grassland, wet meadows, marshes, man-made road verges and railway embankments. Common-spotted Orchids can be found in flower from mid-May to early August.

Pyramidal Orchid (Anacamptis pyramidalis)

Identification: This species is named for the pyramid shape of its flower spike during early growth stages, before it matures into an elongated, cylindrical shape. The flower spike grows up to 60cm in height and is densely flowered with up to 100 flowers at a time. These can be pale pink, to reddish pink in colour and have a sweet vanilla-like scent. The lower lip is divided into three lobes, which can vary in shape, and the sepals and upper petals form a tight hood over the flower. Pyramidal Orchids have up to four grey-green, narrow basal leaves.

Distribution: Common and widespread in England and Wales. It can be found on grassland and sand dunes on chalk and limestone soils, and grows well on human-made habitats, including road verges and roundabouts.

Bee Orchid (Ophrys apifera)

Identification: These distinctive plants have a flower that resembles the body of a bumblebee – a pattern that has since lost significance in the UK, so this species relies on self-pollination.

It grows between 10-50cm in height, and each plant can develop up to seven good-sized flowers. At the base of the stem, there is a rosette of five to six grey green basal leaves – these sit at ground level and have two sheathing leaves that grow upwards on the stem. The distinctive, velvety lip has three lobes – the central section is slightly convex and the side lobes form two small humps. It is rich maroon to purple-brown in colour, with grey and pale-yellow markings. The three wing-like sepals are pink and have a central green vein. Upper petals are brown and cylindrical in shape, forming the antennae of the bee.

Distribution: Found throughout Britain, although scarce in Cornwall, north Devon and Scotland. Bee Orchids can be found on dry chalk and limestone grasslands, calcareous dunes and roadsides or waste ground. Flowers from early June to late July.

Fly Orchid (Ophrys insectifera)

Identification: This fascinating species has evolved to resemble an insect in order to attract pollinators. Fly Orchids can grow up to 60cm in height and can bear up to 15 flowers on a spindly stem. The flower has a velvety texture and is dark purplish brown in colour with an iridescent blue band in the middle of the lip. The lip is long, with two short side lobes, and a central lobe that is notched at the bottom. At the base, there are two coloured patches that resemble eyes, with the same velvety texture. The flower has three sepals that are pointed, yellow green in colour and stiff. The thin upper petals are dark purple or brown and resemble antennae.

Fly Orchids typically have three to four dark green basal leaves, that are shiny on the surface and have blunt tips.

Distribution: Widespread throughout the UK, but populations are declining and are fairly scattered in northern England and north Wales due to habitat loss. Fly Orchids can be found in deciduous open woodland, open scrub, grassland, quarries and roadside banks on chalk and limestone soils, and flowers from May to early June.

Early Purple Orchid (Orchis mascula)

Identification: A distinctive purple flower on a tall spike. Early Purple Orchids can grow up to 60cm in height, and are easily recognisable by their pink-purple flowers and spotted leaves. The flower spike is oval or cylindrical, carries up to fifty flowers at once on a dense cone-shaped cluster, and has darkly spotted, glossy basal leaves that form a rosette on the ground.

Their sepals spread upwards, and the lip is broad with three lobes and lightly crinkled edges. The centre of this broad lip is paler in colour, and features darker coloured spots over the surface. Upper petals and sepals form a hood at the top of the flower, which has a sweet honey-like scent, that quickly turns unpleasant (likened to tomcat urine) as the flower matures. Flowers do not produce nectar and are pollinated by a variety of bees.

Distribution: Early Purple Orchids can be found throughout the UK, with very scattered groups or complete absence in some regions due to habitat loss and agricultural management. These orchids are found mainly on chalk and limestone soils, in woodland, grassland, rocky cliffs, road verges and embankments. Flowers first appear in spring, typically from early April to early June.

Common Twayblade (Neottia ovata)

Identification: Common Twayblades can be readily identified by a pair of broad, round or egg-shaped basal leaves that are flush to the ground. The hairy flower spike can grow up to 50cm in height and contains up to 100 clustered, green flowers that are likened to small people – the upper petals and sepals form a hood over a long, yellowish-green forked lip which resemble legs. They have a slightly musky scent and are pollinated by small insects including wasps, beetles and sawflies.

Distribution: Common and widely distributed throughout the UK. Common Twayblades can be found on grassland, deciduous woodland, dune slacks, road verges and chalky soils. Flowers from late April to early August.

Recommended Reading:

Britain’s Orchids | September 2020

A magnificent work combining close to 100 illustrative plates with over 1,000 colour photos to cover all orchid species, subspecies and varieties, as well as hybrids, at all stages of development.

Orchids of Europe, North Africa and the Middle East | January 2025

Presents the first comprehensive overview of all named orchid taxa from Europe, North Africa and the Middle East, covering etymology, flowering period, habitat, geographical distribution and more for each taxon.

Guide to Orchids | January 2008

With colour paintings covering 69 species, subspecies and hybrids, this 12-panel chart covers all the orchids that are likely to be seen in Great Britain (including the Channel Islands) and Ireland.

Author interview with Jon Stokes: Trees of Britain and Ireland

This comprehensive guide to tree identification features over 3,000 photos and illustrations of more than 300 native and common non-native species. It includes detailed keys, distribution maps and seasonal charts, alongside ecological insights, habitat information and conservation advice on each species, making it an invaluable resource for those interested in British and Irish tree species.

Jon Stokes is one of Britain’s leading tree conservationists and has been studying the world of trees for over thirty years. He is the Director of Trees, Science and Research at a UK based charity The Tree Council, which aims to bring everyone together for the love of trees, and has authored or co-author ten books focusing on trees.

We recently spoke to Jon about Trees of Britain and Ireland, where he told us how he decided which species to include in this book, what his hopes are for the future conservation of our native trees and more.

How did you first become interested in tree biology and ecology, and why did you decide to write the latest WILDGuides identification book?

I have always been fascinated by trees. I love their huge size, their great age and their amazing ability to live for centuries in one spot, coping with everything the world throws at them. To truly understand trees, however, I believe it is vital to understand the other species that depend on them. Oaks, for example, support more than 2,000 other species.

When the opportunity arose to write the WILDGuides tree book, it felt like an amazing opportunity to produce something new – a book that not only allowed the identification of all our diverse and varied native trees and shrubs, but one which also described the ecology of the trees and some of the species which live within them.

What was the greatest challenge you faced when writing this guide, and did you have any specific issues in relation to seasonality?

One of the many challenges in producing this book was our desire to photograph flowers, leaves, fruit and winter twigs – to show how to identify trees all year round. This obviously meant returning to each species a number of times, in all seasons, in all weathers, across the length and breadth of the country. We also wanted to show trees in their natural environments – all of which just made the project a little more complex! Despite the challenges, it was a joyful journey of discovery which took me from the top of Ben Lawers in Scotland; to sea level in Cornwall and Ireland; from sand dunes to heathland; from dense woodland to ancient wood pasture, and occasionally to some very scary cliff edges, looking for Whitebeams.

Trees of Britain and Ireland provides a comprehensive overview of the interdependent relationships between trees and a variety of plant, animal, fungal, and lichen species. Why did you think it was so important to highlight these relationships, and how did you decide which additional species to include?

The web of life that surrounds our trees is truly astonishing, from tiny wasps to bats, from fungi to Ospreys – they all live in and on trees. Selecting the range of species to illustrate was difficult and, of course, there was also a bit of personal preference in the final choices! In some cases, the selection process was very difficult. To my surprise, the biodiversity of some species, like Wild Cherry, doesn’t appear to be well studied – something I hadn’t realised until working on this book.

It was evident throughout the book that this is a clear, easily digestible guide suitable for readers of all abilities. How important do you think nature accessibility and education is for future generations?

I have had the pleasure of spending my working life engaging people with nature. From guided walks, to lectures and talks, I love showing people new aspects of the world around us. Life is fascinating and every day is a school day. For example: Why are leaves green? How do trees grow? Why is that tree, that shape? These are the questions that enthral me daily. I guess this is a long-winded way of saying that I believe learning about nature is vital to all of us, all the time, and at any age. I hope this book sparks an interest in learning more about my beloved trees. I hope it sparks a desire to go out and explore other aspects of our beautiful islands and the wealth of wildlife we have here.

What patterns did you notice whilst researching conservation status in the UK, and what are your hopes for the future conservation and protection of our native trees and plants?

This is a really interesting question. I now realise that, before researching this book, I hadn’t really thought about the conservation status of many of our trees and shrubs. Yet, Britain and Ireland actually have many rare and globally endangered species growing exclusively here, like many of the endemic Whitebeams and the Wild Cotoneaster. Our knowledge of our rare trees is sometimes scant. In the last twelve months, new information has emerged about our elm trees, suggesting there may be new and rare elm species hiding in plain sight.

So, what are my hopes for the future of these rare species? Well, if nothing else, the development of the book allowed me to clearly articulate the importance of these rare trees in my work. More importantly, I hope a better understanding of the importance of these rare trees will allow better protection to be developed for these globally rare species growing in these islands.

What’s taking up your time at the moment? Are you working on any other books or projects that we can hear about?

In terms of projects, there is one current Tree Council project that is very important to me. This project draws on work undertaken twenty years ago when, with my fellow authors and photographers, we had the privilege of writing a number of books on our ‘Heritage Trees’ – trees that are as important to the nation as some of our great stately homes and castles. From 2000-year-old yews, to astonishing old broadleaved trees like the Bowthorpe Oak, we featured a range of these iconic trees to show the amazing living heritage we have in Britain and Ireland. Now 20 years on, we are revisiting the trees to see how they have fared over the last two decades and the results of this will be published later in the summer. It’s already clear that while some of the trees have thrived over this time, others have not done so well. Our aim is for this project to lead to the development of new guidance on how to better protect our most important trees, to ensure that we can marvel at these ‘Green Monuments’ and hear their stories for many decades to come.

Trees of Britain and Ireland is available from our online bookstore.

Author interview with Richard Lewington: Pocket Guide to British Spiders

Featuring 130 of the most common and readily available spider species, Pocket Guide to British Spiders is the ideal companion for both beginners and more experienced enthusiasts alike. Detailing identifying features, webs, egg cocoons and behaviour, these accounts are interspersed with beautifully illustrated spreads showing similarities and differences at a glance, making this introductory guide the perfect companion for use in the field.

Richard Lewington is a renowned wildlife illustrator, whose beautifully detailed drawings feature in a wide array of identification guides including The Butterflies of Britain and Ireland, Field Guide to the Moths of Great Britain and Ireland, and Field Guide to the Bees of Great Britain and Ireland. In 1999, he was awarded Butterfly Conservation’s Marsh Award for the promotion of Lepidoptera conservation, as well as the Zoological Society of London’s Stamford Raffles Award for contribution to zoology in 2010.

We recently had the opportunity to speak to Richard about his latest book, including how he first became an author, why he chose to focus on arachnids and which techniques he uses to create these stunning illustrations.

Firstly, can you tell us a little bit about yourself, and how you first became an author and illustrator?

I became a freelance wildlife illustrator soon after leaving art college in the 1970s. I’ve always had an interest in the natural world, particularly butterflies and moths, so invertebrates were the subjects I concentrated on right from the start. I’ve always worked in collaboration with various experts, until about 25 years ago, when I wrote and illustrated my first book How to Identify Butterflies. Since then, I’ve written and illustrated five books, though most of the books I work on are collaborations with authors with specialist knowledge of the subjects.

Pocket Guide to British Spiders is your 29th identification guide published since 1979. What inspired you to focus on arachnids for this book, and how did you choose which of the 680 British species to include?

I’ve had an interest in spiders for many years and have a collection of paintings, some of which I did about 50 years ago. I thought they would form the basis of a pocket guide, and I added more illustrations in the last few years to go with those I already had. The aim was to include common, well-known and easily identifiable species – about 130 in total. As many spiders are very small and similar to each other, it is necessary to examine them through a microscope to identify many of them, which is the next step up for those who want to study them in greater detail.

I was surprised to learn about the somewhat limited conservation efforts being undertaken to protect endangered spider species across Britain. What are the largest threats to spiders, and how can we protect these vital invertebrates in the future?

As with much of the natural world, spiders are under threat from habitat loss and pollution, but because they don’t have the same appeal as some other groups, they receive less attention. However, they are just as important, helping to keep the balance in ecological systems by eating and being eaten by a vast number of other creatures. Gardens are becoming more important, and we can all help spider well-being by not being too tidy in the garden to encourage a good variety of mini habitats in which they can live.

Pocket Guide to British Spiders includes a vast amount of detail on nomenclature, structure, anatomy, mythology, identifying features and more. What was the most interesting thing you learned while researching this book?

Probably their diversity. They vary hugely in size, shape and colouring, and can be found in every habitat from mountain tops to beneath the surface of water. Many also occur throughout the year when many other invertebrates are dormant.

Having illustrated many guides throughout your career, can you give us an overview of the techniques you use to create such detailed works of art, and explain how your painting style has evolved over time?

I start by doing a detailed, measured drawing on layout paper. This is transferred to watercolour paper and, using Designers’ Gouache paints, I apply washes, gradually building the colour intensity and the effect of light and shadow. Details such as textures and highlights are added to give a three-dimensional appearance. More recently, I’ve been scanning the artwork before sending it to the publishers, so that I can adjust, enhance and touch-up the images using a tablet and image enhancing software.

What were the most difficult aspects of illustrating spiders, given their intricate and often microscopic features?

Spiders can’t be preserved in museum collections like butterflies and beetles, as they have soft bodies which shrivel after death, so working from live specimens or from detailed photographs are the best options. This means going into the countryside to find specimens and asking other spider enthusiasts to help. I like to take my own macro photographs from the exact angle to avoid any distortion, I can then work using the photos and the live specimen, observing it under my microscope. It’s also necessary to have a good knowledge, so that the subjects can be correctly identified, as some are very similar.

Finally, what’s taking up your time at the moment? Are you planning to write or illustrate any more books over the coming years?

My next book will be a field guide to Orthoptera, which I completed several years ago but has been long delayed. After that I shall be working on a project about my work as a wildlife illustrator, concentrating mainly on British moths, butterflies and their caterpillars.

Pocket Guide to British Spiders is available from our online bookstore.

The NHBS Guide to UK Grassland Fungi

In the past 75 years, over 90% of western Europe’s grasslands have disappeared. These overlooked and undervalued habitats deliver essential ecosystem services that contribute to the function of the planet – they provide key wildlife habitat, can store large amounts of carbon, and are instrumental in mitigating soil erosion and flood management, which is essential as our landscapes adapt to climate change.

Grassland fungi – such as those outlined in this identification guide – form a distinctive community in these habitats and are key indicators of ancient meadow and unimproved grassland. These species fruit from late summer until winter, and vary greatly in size, shape, colour and texture, from the pale, spherical Earthball to vibrant, slender coral fungi.

In this guide, explore a selection of grassland fungi groups, as well as individual species, that are commonly found across Britain, and read about their identifying features, size and distribution.

Groups

Waxcaps (Hygrocybe)

Identification: There are about 150 different species of Hygrocybe (meaning ‘watery head’) in the UK, and these are a very easily identifiable group. They are characterised by their vibrant, smooth waxy caps that come in a range of bright colours, including red, pink, yellow, orange and brown. When present on the fruiting body, gills are thick but evenly coloured and spaced.

Distribution: Found in nutrient-poor grasslands, such as grazed upland, and often grow alongside other groups including Pinkgills, Corals, Clubs and Earthtongues throughout late summer and autumn.

Size: 5-7cm

Club and Corals (Clavariaceae)

Yellow club fungus. — Yellow club fungus by Jo Graeser.

Identification: These distinctive fungi are easily recognisable by their slender, forking branches that are often rooted in buried wood. They have thick, dense finger or club shaped stalks that grow from a central base, often in clusters, and can be bright yellow, pale pink or white. Coral fungi produce yellow spores from the tip of their branches.

Distribution: Coral fungi predominantly grow on wood, but can also be found in soil, leaf litter or in short grasses and moss from July to November.

Size: 4-10cm

Wood Pinkgill (Entoloma rhodopolium)

Brown caps of wood pinkgill mushrooms (Entoloma rhodopolium) in Huntsville State Park. Texas. — Brown caps of wood pinkgill mushrooms (Entoloma rhodopolium) via Texas Mushrooms.

Identification: Often found in large groups, this very common pinkgill mushroom is defined by its convex, rounded fruiting body that flattens at the bottom of the skirt. Juvenile mushrooms have tightly packed white gills, and turn a light shade of pink with maturity. They feature a long, smooth, lightly coloured stem measuring 4-9cm long.

Distribution: Most commonly found throughout the summer until late autumn in deciduous broadleaf woodlands.

Size: 3-5cm

Note: Poisonous

Species

Earthtongue fungi (Geoglossum fallax)

Earthtongues - Glutinoglossum by Lukas Large, via flickr. — Earthtongues – Glutinoglossum by Lukas Large, via flickr.

Identification: A widespread species found across Britain and Ireland, Earthtongues are characterised by black or dark brown stromata with a single, club-like fruiting body growing from a cylindrical, scaly stem. The head is grooved and makes up a third of the body, and it is covered in fine hairs to protect its fertile spores.

Distribution: Grows in mossy, unimproved and acidic grassland such as old churchyards from August to November.

Size: 2-7cm

Common Inkcap (Coprinopsis atramentaria)

Inkcap Trio by Derek Winterburn, via flickr.

Identification: Starting off as a smooth, egg-shaped ball with scaled central area, the grey or greyish-brown cap later expands into a soft bell shape with a slight curve at its margin. The stem of an inkcap is smooth and reddish-brown in colour, with hundreds of crowded gills that are white at first, but later darken to brown and black before being auto-digested by the fruiting head.

Distribution: Fruits in small clumps on tree stumps, buried hardwood, woodland footpaths and parkland from May until November. Some solitary specimens have been sighted on occasion.

Size: 3-7cm

Note: Poisonous

Petticoat Mottlegill (Panaeolus papilionaceus)

Identification: Petticoat Mottlegills are recognisable by their smooth bell caps with a serrated edge, and can be pale brown, greyish-brown or pallid grey, with a darker centre. The fused-together gills are a pale, greyish brown with white edges that turn dark brown and black with maturity, and the thin, cylindrical stem is covered in a fine white powder.

Distribution: Commonly located in groups on rotted dung from June to November.

Size: 6-12cm

Common Earthball (Scleroderma citrinum)

Identification: Ranging from light ochre to mid-brown or green, with a yellow tinge on its upper surface, the Earthballs rounded, fruiting body is attached to the ground with white mycelial threads and is stemless. It has tough, thick skin covered in coarse scales that vary in size and shape, which rupture as the head matures to allow the wind and rain to disperse its spores – which are white when young, and darken to brown or purple with age. These empty shells often remain in sheltered woodland hollows for many months before decaying.

Distribution: Acidic, well-drained soil near forest paths or on shady banks from July to early December.

Size: 4-10cm

Common Parasol (Macrolepiota procera)

Parasol Fungi by Bernard Spragg, via flickr.

Identification: This edible species of mushroom is round in shape with a pale brown, spherical cap that darkens as it nears the crown. The cap breaks into a scaly texture that features a darker brown, central bump called an umbo. The broad, crowded gills of Common Parasol mushrooms are white or pale cream, and the cap has a white flesh when cut into. The tough, fibrous stem is surrounded by a smooth, white double-edged ring decorated with small brown scales that are comparable to snakeskin.

Distribution: Commonly found on verges, neglected grassland or cliffs, with multiple mushrooms growing in a slightly wavy line, from July to November.

Size: 10-25cm

The NHBS Guide to UK Bracket Fungi

Bracket fungi (basidiomycetes), also known as shelf or polypore fungi, can grow on both dead or living trees and cause widespread heartwood decay deep inside its host. These fungal bodies appear year-round depending on the species and produce bracket-shaped fruit on the trunk, roots or branches of the host tree that can span up to 70cm in diameter.

There are over 1,000 species of bracket fungi worldwide, and here you can explore a selection of bracket fungi found in Britain, detailing their identifying features, when and where they can be found.

Chicken of the Woods (Laetiporus sulphureus) – also known as Sulphur polypore’

Identification: This easy to spot, sulphur-yellow fungus can grow up to 40cm in diameter and features numerous thick, soft, overlapping brackets. When young, they have a velvety texture and orange bands across yellow flesh, which fades to light cream with age. Each bracket is fan-shaped with an undulating margin, while the underside has small pores and produces a yellow liquid when squeezed.

Distribution: June to November. Mainly grows on oak tree trunks, but can also be found on Yew, cherry, chestnut and willows.

Size: 10-40cm

Hen-of-the-Woods (Grifola frondosa)

Identification: This large, very distinctive fungus features many flat, interlocking lobes growing from a dense centre attached to the foot of a tree. The underside of the fronds are cream in colour, their top side can vary from brown to grey, and the flesh is pale cream. It’s very common for multiple growths to emerge from one host, and fully grown hen-of-the-woods can weigh several kilograms.

Distribution: August to November. Predominantly around the base of mature oak trees but they do occasionally grow on other deciduous species.

Size: Up to 70cm

Turkeytail (Trametes versicolor)

Turkeytail Fungi (Trametes versicolor) by Jo Graeser.

Identification: Turkeytail fungi produce thin but tough semi-circular caps in tight, overlapping tiers that have a slightly hairy, velvety texture. It has a range of concentrated coloured rings that are a mix of black, brown, grey, yellow, purple and green, however the outer ring along the edge of the bell is pale in colour. Often confused with Stereum ostrea (or False Turkeytail) this species can be correctly identified by its white underside featuring hundreds of visible pores.

Distribution: Can be found on fallen hardwood trees such as Beech or oak throughout the year, but is most abundant from late spring to autumn.

Size: 4-10cm

Beefsteak (Fistulina hepatica)

Beefsteak fungus (WGP). — Image by David Short via flickr.

Identification: This bracket fungus can be pink, red or brown in colour, with a light underside full of pores that turn red with age. Featuring a slightly inflated edge that flattens as it matures, Beefsteak fungus is moist to the touch and produces smooth, round, pale pink spores.

Distribution: August to November. Usually found in broadleaved woodlands on oak trees.

Size: 6-25cm

Birch Polypore (Piptoporus betulinus) – Also known as the Razorstrop fungus

Identification: Birch Polypore is a common bracket fungus that grows exclusively on birch trees. It has a rounded, brown-white cap with a rubber or leather-like texture and white pores on its underside.

Distribution: It can be found year-round on the lower trunk of birch trees and fallen birch branches.

Size: 10-20cm

Dryads Saddle (Cerioporus squamosus)

More Dryad's saddle. — Image by Andrew Hill via flickr.

Identification: This species of fungi grows in overlapping tiers on dead or dying broad-leaved trees, particularly elm, Beech and Sycamore. Recognisable by their distinctly textured, creamy-ochre brackets, these fungiemerge at soil level at the base of the host tree or higher up the trunk. The underside of the fruit features hundreds of honeycomb-shaped pores.

Distribution: This fungus fruits in the summer and again in autumn, and is most commonly seen from April to September on deciduous trees and fallen wood on the forest floor.

Size: Up to 70cm

Southern Bracket (Ganoderma australe)

Identification: Southern Bracket fungi can grow on both living and dying deciduous trees and first appear in the form of solid, white balls. Over a number of years, these develop into large, thick brackets with a flat brownish top and a white underside. As the fungus ages, the smooth top can become ridged and bumpy, and when the underside is scored it produces dark brown marks. This fungus can be easily confused with the Artists Fungi ( applanatum), however they can be distinguished by examining their spores under a microscope.

Distribution: This is the most common species of the Ganoderma genus and fruits on most deciduous trees from April to August.

Size: Up to 60cm

King Alfred’s Cakes (Daldinia concentrica)

Identification: These fungi present themselves in the form of hard, round lumps roughly the size of a golf ball and are a matt, pinky-brown colour when young. As they mature, they darken to a coal colour and become shiny in appearance. The outer surface is solid but cracks easily, while the inside of the fruit is easily identified by black and grey concentric rings that are darker than the flesh. King Alfred’s Cakes are also known as coal fungus as they can be used as kindling or firelighters.

Distribution: Can be found year-round in deciduous woodland, especially on fallen Beech and Ash branches.

Size: 2-10cm

Horses Hoof (Fomes fomentarius)

Horse Hoof Fungi, Knole Park, Sevenoaks, Kent. — Image by Nigel Turner via flickr.

Identification: As evident from the name, this species of fungi is typically shaped like a horse’s hoof, with a pointed top and wide bottom. Featuring concentrated ridges across its exterior and with a fleshy, fibrous inside, their blunt, rounded margin is often cream or yellow. Although they can vary from silvery-grey to black in colour, most specimens are shades of brown. Small circular pores under the bell start off cream in colour and darken to brown once the fruit has matured. On occasion, some Horses Hoofs appear in more of a traditional, flat bracket shape.

Distribution: Can be found growing year-round on weak, living or dead hardwood deciduous trees, particularly Birch.

Size: 5 to 45cm

The NHBS Guide to Grasshoppers and Bush-crickets

Here in the UK, we have 11 species of grasshopper and 23 species of cricket (around ten of these being bush-crickets), both subfamilies belonging to the order Orthoptera. Although similar in appearance, the two can be separated by their antennae – grasshoppers have short, stout appendages and bush-crickets have longer, thinner antennae. Bush-crickets also have a larger body size than grasshoppers, growing up to 5cm in length, and females have a distinctive ovipositor used to lay eggs.

Here, we look at a few species of grasshoppers and bush-crickets found in the UK, detailing ways to ID them and where they can be found.

Common Field Grasshopper (Chorthippus brunneus)

A brown grasshopper is sat on a small twig, it is dull in colour but has a vibrant yellow-orange abdomen under a darker wing — Image by Gilles San Martin via Flickr

ID Notes: Up to 2.5cm in length. This grasshopper is usually a mottled brown colour, but can vary from brown to grey, orange and purple. It has barring on the sides, and a very hairy underside which is the most straightforward way to identify this species. It is winged and able to fly. Their song consists of short, single chirrups repeated at short intervals.

Distribution: Common and widespread throughout Britain. Common Field Grasshoppers can be seen from May to October on short vegetation in dry, warm days. They can be found in grassy areas including farmland, grassland, heathland, moorland, towns and gardens.

Meadow Grasshopper (Chorthippus parallelus)

A grasshopper on a green stem, it has a vibrant pink head, thorax and back legs. It has a green underbelly and black/yellow striping on its abdomen. Its wings are very short. — Image by Gilles San Martin via Flickr

ID Notes: Up to 2.3cm in length. Meadow Grasshoppers are usually green in colour, with some individuals brown or pink-purple coloured (pictured above). The wings are often brown and are long in males, whereas female wings are short and do not reach the end of the abdomen. This species has distinctive black knees and a dark brown stripe running across the flank to the eye. They are similar in appearance to Common Field Grasshoppers, but lack a hairy underside, which can be used to distinguish them from one another. Their song is a burst of around ten chirrups, making a ‘rrr’ sound.

Distribution: Common and widespread throughout Britain. Meadow Grasshoppers can be seen from April to October in damp pastures with long grass. They can also be found in meadows, grassland, heathland and moorland.

Common Green Grasshopper (Omocestus viridulus)

A green grasshopper is standing on a bed of moss, it is light green in colour with dark brown striping on its abdomen — Image by Billy Lindblom via Flickr

ID Notes: Up to 2.3cm in length. As the name suggests, this species is mostly green in colour, but males can also be olive brown. Lines on the shoulders are incurved and white or cream in colour. Common Green Grasshoppers are winged and can fly, but their wings do not exceed the body’s length. They produce a long, loud ticking song which can last for 20 seconds or more and is reminiscent of a fast-paced freewheeling bicycle.

Distribution: Common and widespread throughout Britain, although a little patchier in the lowlands. This species appears early in spring and can be seen from April to September. It can be found in coarse, moist grass in meadows, woodland rides, hillside pastures and farmland.

Mottled Grasshopper (Myrmeleotettix maculatus)

a brown and green grasshopper. it has small antennae a green head and green patches on the thorax and rear legs. the rest of the body is mottled brown in colour — Image by Frank Vassen via Flickr

ID Notes: Up to 2cm in length. This small grasshopper is most easily identified by its antennae – males’ are club-tipped, and females’ have a thickened tip. The body has several colour variations including green, brown, grey and orange. Their song is a repetitive ‘zrr’ ‘zrr’ sound lasting around ten seconds before stopping abruptly.

Distribution: Widely distributed across the UK, but uncommon. Mottled Grasshoppers can be found from April to October in areas of dry, short grass with access to rocks or bare ground, including grasslands, heathland and old quarries.

Oak Bush Cricket (Meconema thalassinum)

A small, lime green cricket is sat on three leaflets. It has very long, flexible antennae that are trailing off of the leaf and long, yellow legs — Image by Alison Day via Flickr

ID Notes: Up to 1.7cm in length. This species is the UK’s only native, arboreal cricket. It has a slender, lime green body with a yellow-orange or brown strip running along its back. Both sexes are fully winged, with wings reaching the end of the abdomen. Females have a long, upturned ovipositor and males have a pair of shorter, rounded claspers. The males do not produce a song, but instead drum their hind legs on nearby leaves.

Distribution: Common, found in England and Wales. The Oak Bush Cricket is common in the midlands and the south but is absent further north. It can be found in the canopy of mature trees in woods, hedges, garden shrubs and other deciduous trees from June to November.

Dark Bush Cricket (Pholidoptera griseoaptera)

A dark brown cricket is standing on a large leaf. It has very long thin antennae and a curved, pointed spine at the end of its body. Its long rear legs are outstretched, standing on another leaf — Image by Alison Day via Flickr

ID Notes: Up to 2.1cm in length. This species is typically reddish to dark brown, with occasional yellow green through the abdomen and a paler area along the top of the thorax. It has very small forewings and long, hair-like antennae. Females have a large, upcurved ovipositor.

Distribution: Fairly common in central and southern England and Wales, where it can be found in woodland, hedgerows, grassland, farmland, towns and gardens. It can often be spotted in large numbers around bramble patches, where they are basking in the sun.

Roesel’s Bush Cricket (Metrioptera roeselii)

A brown and green cricket with long rear legs and an upturned hook at the end of the abdomen. — Image by Charlie Jackson via Flickr

ID Notes: A short-winged and usually flightless species, Roesel’s Bush Crickets are typically dark brown or yellow in colour, sometimes tinged green. They have a green face, slightly orange legs and a cream-coloured margin to their thorax, with two or three yellow spots on the side. Their monotonous song is recognisable for its long, mechanical sound.

Distribution: Common, found mostly in the southeast of the UK, but currently expanding its range north and west. This species can be found from June to October in rough grassland, scrub, salt marshes and damp meadows.

Great Green Bush Cricket (Tettigonia viridissima)

A large green cricket is resting on a leaf. It has long, slightly yellow legs and its green wings extend further than the abdomen — Image by Gunera via Flickr

ID Notes: This large bush-cricket can easily be distinguished by its size, growing up to 7cm long. It is a vibrant green with an orange-brown stripe running the length of its back. Their wings are typically longer than the abdomen, and females have a long, downward curved ovipositor. This species has a loud, long song which sounds like a high-pitched sewing machine running continuously.

Distribution: Common throughout south England and Wales. It can be found from May to October in scrub, woodland, grassland and heathland, with a preference for light, dry soils.

Further Reading
Grasshoppers of Britain and Western Europe

Paperback | June 2021

The first guide to the 261 species of orthopterans of Britain and western Europe ever published. Features stunning, detailed images and information on the distribution and identification criteria. Includes a CD with over 200 orthopteran songs to aid identification.

Orthoptera and Dermaptera

Paperback | July 2024

A photo-based identification guide covering 28 native breeding species of the UK. This guide is part of FSC’s AIDGAP series (Aids to Identification in Difficult Groups of Animals and Plants), providing accessible identification tools for naturalists over the age of sixteen.

Guide to British Grasshoppers and Allied Insects

Unbound | August 1999

A reliable, fold-out chart with an introduction and a key to British grasshoppers and related insects, ideal for carrying in-field.

The NHBS Guide to UK Centipedes and Millipedes

If you look under logs, stones or through piles of leaf litter, you may find a Myriapod (a type of arthropod with many legs, related to insects and spiders). Although these misunderstood animals may strike fear in some, these invertebrates play a vital role in maintaining our ecosystems. One of their main roles lies in nutrient cycling, where they feed on organic matter and detritus, breaking materials down into a simpler form and thus releasing nutrition back into the soil. They also play an important role, both as predators and prey, in the ecosystem’s food chain and are often celebrated for their effectiveness as a natural biocontrol agent.

The UK has more than 50 species of centipede. This group are generally carnivorous and have around fifteen pairs of legs, with one pair present on each body segment. They have large forcipules, a pair of pincer-like, modified front legs that are used to inject venom into their prey. There are around 65 species of millipede in the UK, which can be distinguished by their shorter, more clubbed antennae, the absence of forcipules and their tendency to have two pairs of legs on most body segments.

Here, we look at a few species of centipede and millipede that can be found in the UK, detailing ways to ID them and where they can be found.

Common Centipede (Lithobius forficatus)

A small orange-brown centipede on a piece of wood, curled up in a c shape — Image by Frank Vassen via Flickr

ID Notes: Also known as the Brown Centipede, Lithobius forficatus is one the largest centipede species in the UK, growing up to 3cm in length. They have long, thin segmented bodies that are a reddish-chestnut brown colour. Each segment has one pair of legs, with 15 pairs in total. They have long antennae and an elongated pair of back legs, giving the illusion of a second pair of antennae at the rear of the body. This species is best identified by thorned projections on the outer edges of the 9th, 11th and 13th segments.

Distribution: Very common and widespread across the UK. This species can be found year-round but is most numerous in spring and autumn. They can be found in a variety of habitats, including woodland, grassland and gardens, and will spend most of the day nesting in soil or sheltered areas under stones, bark or dead wood before emerging at night to hunt for prey.

Stone Centipede (Lithobius microps)

an orangey brown centipede with many legs on a rock — Image by Keith Lugg, the British Myriapod and Isopod Group

ID Notes: Similar in colour to the Common Centipede, Lithobius microps is a ground-dwelling species that is usually chestnut brown in colour, although red and orange varieties can also be found. It is much smaller than its common cousin, reaching around 1cm in length, and its 15 pairs of legs are slightly shorter and paler than those of the Common Centipede.

Distribution: Fairly common in the south of Britain. This fast-moving species can be found year-round in gardens and bare soil in rotting wood and under rocks and stones.

Banded Centipede (Lithobius variegatus)

A brown centipede with striped legs is crawling over a stick on muddy floor, it has many legs and a large head — Image by J. P. Richards, the British Myriapod and Isopod Group

ID Notes: Measuring up to 3cm in length, this small, yellow-brown centipede is easily distinguished by the banding on its legs – these alternate pale and dark, are usually brown in colour and are more obvious on the rear pairs. It has a dark band running centrally along its back and much larger jaws when compared to other Lithobius species.

Distribution: Widespread and common in the UK. It can be found year-round living under stones and decaying plant matter during the day, and is commonly found in garden compost heaps.

Flat-backed Millipede (Polydesmus angustus)

a flat, armoured millipede with many legs and segments walking along muddy ground — Image by J. P. Richards, the British Myriapod and Isopod Group

ID Notes: This millipede grows to around 2.5cm in length, with a width of 4mm. It has a long, flat body with around 20 body segments, each containing between one and two pairs of legs. It is orange-brown in colour and has numerous off-white coloured legs.

Distribution: Widespread and common in the UK. They can be found in almost any environment in which millipedes occur, with frequent sightings in woodland under dead wood and leaf litter.

Striped Millipede (Ommatoiulus sabulosus)

a brown millipede with two orange stripes running down its body on its back. it is walking across a tree stump — Image by S. Rae via Flickr

ID Notes: Up to 3cm in length, this species is one of the largest millipedes found in Britain. It is thick and cylindrical in shape, usually dark brown to black in colour. It has orange coloured stripes running the length of its body on its back, but these can sometimes appear as a series of orange patches.

Distribution: Common and widespread throughout Britain and Ireland. The Striped Millipede can be found year-round in meadows, fields and woodland under leaf litter and logs. It is also known to climb on walls and trees to feed on algae and can be found during the day.

White-legged Snake Millipede (Tachypodoiulus niger)

A curled up black millipede, with paler almost white legs — Image by AJ Cann via Flickr

ID Notes: The White-legged Snake Millipede can grow to around 6cm in length. It is tubular-shaped with a shiny black body and contrasting white legs, of which there are around 100. Immature millipedes are brown, often with pale, longitudinal stripes down the back. It has a pointed telson (legless tail segment) at the end of its body.

Distribution: Common and widespread in the UK. This species can be found year-round in most habitats containing millipedes, including woodland and gardens, or areas with rocks or rotting vegetation to provide cover.

Additional Reading

Atlas of the Centipedes of Britain and Ireland

Paperback | January 2023

A comprehensive atlas of the 55 species of centipede in Britain and Ireland,. tThis guide provides species accounts, distribution maps and detailed colour photographs for easy identification.

Key to the Identification of British Centipedes

Paperback | December 2008

An identification guide from the Field Studies Council AIDGAP series, this book contains keys to the 57 species of centipede found in Britain.

Common European Yellowjacket (Vespa vulgaris)

Red Wasp (Vespula rufa)

Hornet (Vespa crabro)

Yellow-legged Asian Hornet (Vespa velutina)